Abstract
Major histocompatibility complex (MHC) molecules play an important role in antigen presentation for induction of tumor as well as cellular and humoral immunities. Recent studies using anti-MHC antibodies demonstrated that antibodies specific for HLA class I molecules induced cellular activation and a type of apoptosis that may be distinct from Fas-dependent or TNFR (tumor necrosis factor-alpha receptor)-dependent processes. We purified a previously untested apoptosis-inducing factor from HL-60 human leukemic cell-conditioned media to homogeneity and sequenced it. It was identified as beta(2)-microglobulin (beta(2)m), which has been previously known as thymotaxin and is a part of the HLA class I antigen complex. beta(2)m acts on both T-leukemic cells and myeloid leukemic cells to induce apoptosis, which then activates caspase 1 and 3. Cross-linking studies showed that biotinilated beta(2)m recognized an epitope distinct from those recognized by the anti-HLA class I antibody, as reported previously. We demonstrated that beta(2)m plays a previously unrecognized and important role in regulating the elimination of tumor cells, which occurs as a result of the action of beta(2)m as an apoptosis-inducing factor.
Publication types
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Comparative Study
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Research Support, Non-U.S. Gov't
MeSH terms
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Amino Acid Chloromethyl Ketones / pharmacology
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Amino Acid Sequence
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Animals
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Antibodies, Monoclonal / pharmacology
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Apoptosis / drug effects*
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Culture Media, Conditioned / chemistry
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Cysteine Proteinase Inhibitors / pharmacology
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Fas Ligand Protein
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HL-60 Cells / chemistry
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Humans
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K562 Cells / drug effects
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Membrane Glycoproteins / physiology
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Mice
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Molecular Sequence Data
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Neoplasm Proteins / immunology
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Neoplasm Proteins / isolation & purification
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Neoplasm Proteins / pharmacology
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Neoplasm Proteins / physiology
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Oligopeptides / pharmacology
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Receptors, Tumor Necrosis Factor / physiology
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Sequence Alignment
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Sequence Homology, Amino Acid
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Tumor Cells, Cultured / drug effects
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Tumor Necrosis Factor-alpha / physiology
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U937 Cells / drug effects
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beta 2-Microglobulin / immunology
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beta 2-Microglobulin / isolation & purification
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beta 2-Microglobulin / pharmacology*
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beta 2-Microglobulin / physiology
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fas Receptor / physiology
Substances
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Amino Acid Chloromethyl Ketones
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Antibodies, Monoclonal
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Culture Media, Conditioned
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Cysteine Proteinase Inhibitors
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FASLG protein, human
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Fas Ligand Protein
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Fasl protein, mouse
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Membrane Glycoproteins
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Neoplasm Proteins
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Oligopeptides
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Receptors, Tumor Necrosis Factor
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Tumor Necrosis Factor-alpha
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benzyloxycarbonylvalyl-alanyl-aspartyl fluoromethyl ketone
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beta 2-Microglobulin
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fas Receptor
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L 709049