Adenine nucleotide N-glycosidase activity of the A-chain of cinnamomin characterized by 1H-nuclear magnetic resonance

Biol Chem. 2000 May-Jun;381(5-6):447-51. doi: 10.1515/BC.2000.058.

Abstract

Plant ribosome-inactivating proteins specifically cleave an N-glycosidic bond of a unique adenosine in the largest ribosomal RNA, releasing an adenine from ribosomes of different sources. Here, 1H-nuclear magnetic resonance is used to analyze the enzymatic products of the A-chain of cinnamomin, a type-II ribosome-inactivating protein (RIP) acting on the nucleotides in situ. The enzymatic activities of the RIP on nine nucleotides are compared. Cinnamomin A-chain can cleave the N-glycosidic bond and release an adenine base from adenine nucleotides except 5'-ATP; however, it cannot act on 5'-GMP, 5'-CMP, and 5'-UMP. The A-chain in the mixture of cinnamomin A- and B-chain exhibits higher activity toward adenine nucleotides than the A-chain alone does, suggesting that the B-chain can conformationally stabilize the A-chain. Intact cinnamomin also exhibits lower activity toward adenine nucleotides. However, cinnamomin B-chain and heat-denatured intact cinnamomin cannot hydrolyze all the tested nucleotides. We conclude that hydrolysis of the N-C glycosidic bond of nucleotide compounds by cinnamomin A-chain has a base preference, and the negatively charged phosphate group(s) reduces the recognition ability of the A-chain to adenine nucleotide.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adenine Nucleotides / metabolism*
  • Algal Proteins
  • Glycoside Hydrolases / metabolism*
  • Molecular Weight
  • Nuclear Magnetic Resonance, Biomolecular
  • Proteins / chemistry
  • Proteins / metabolism*
  • Ribosome Inactivating Proteins, Type 2
  • Substrate Specificity

Substances

  • Adenine Nucleotides
  • Algal Proteins
  • Proteins
  • Ribosome Inactivating Proteins, Type 2
  • cinnamomin, Phytophthora cinnamomi
  • Glycoside Hydrolases