Abstract
Scurfy (sf) is an X-linked recessive mouse mutant resulting in lethality in hemizygous males 16-25 days after birth, and is characterized by overproliferation of CD4+CD8- T lymphocytes, extensive multiorgan infiltration and elevation of numerous cytokines. Similar to animals that lack expression of either Ctla-4 or Tgf-beta, the pathology observed in sf mice seems to result from an inability to properly regulate CD4+CD8- T-cell activity. Here we identify the gene defective in sf mice by combining high-resolution genetic and physical mapping with large-scale sequence analysis. The protein encoded by this gene (designated Foxp3) is a new member of the forkhead/winged-helix family of transcriptional regulators and is highly conserved in humans. In sf mice, a frameshift mutation results in a product lacking the forkhead domain. Genetic complementation demonstrates that the protein product of Foxp3, scurfin, is essential for normal immune homeostasis.
MeSH terms
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Amino Acid Motifs
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Amino Acid Sequence
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Animals
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Cloning, Molecular
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Conserved Sequence
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DNA Mutational Analysis
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DNA-Binding Proteins / chemistry*
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DNA-Binding Proteins / genetics
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DNA-Binding Proteins / metabolism*
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Female
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Forkhead Transcription Factors
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Gene Expression Profiling
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Genes, Essential / genetics*
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Genes, Recessive / genetics
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Genetic Complementation Test
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Humans
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Lymph Nodes / immunology
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Lymph Nodes / pathology
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Lymphocyte Count
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Lymphoproliferative Disorders / genetics*
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Lymphoproliferative Disorders / immunology
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Lymphoproliferative Disorders / pathology
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Male
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Mice
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Mice, Mutant Strains
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Mice, Transgenic
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Molecular Sequence Data
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Mutation / genetics*
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Phenotype
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Physical Chromosome Mapping
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Protein Structure, Tertiary
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RNA, Messenger / analysis
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RNA, Messenger / genetics
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Sequence Alignment
Substances
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DNA-Binding Proteins
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FOXP3 protein, human
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Forkhead Transcription Factors
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Foxp3 protein, mouse
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RNA, Messenger
Associated data
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GENBANK/A49395
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GENBANK/AF196779
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GENBANK/AF235097
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GENBANK/AF277991
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GENBANK/AF277992
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GENBANK/AF277993
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GENBANK/AF277994
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GENBANK/AF277995
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GENBANK/AF277996
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GENBANK/AF318279
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GENBANK/AF318280
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GENBANK/AF318281
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GENBANK/AJ005891
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GENBANK/U93305
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GENBANK/X97571