Abstract
H2-O/HLA-DO are MHC class II accessory molecules that modulate exogenous Ag presentation. Most class II accessory molecules are expressed in all professional APC; however, H2-O is only expressed in B cells and medullary thymic epithelial cells. Because B cells present exogenous Ags and superantigens (SAgs), and medullary thymic epithelial cells are specialized APC for self Ags during negative selection in the thymus, we have hypothesized that H2-O might play a role in MHC class II-restricted SAg and self Ag presentation. In this study, we demonstrate that H2-O expression inhibits presentation of the bacterial SAgs staphylococcal enterotoxins A and B to four SAg-reactive T hybridoma cells. In contrast, H2-O has no effect on presentation of endogenous self Ags, as measured by tumorigenicity in vivo and Ag presentation to three self Ag-specific T hybridoma cells. Additional experiments suggest that H2-O inhibits presentation of exogenous Ags by both newly synthesized and recycling MHC class II molecules. These data suggest H2-O may have a physiological role in tolerance induction and SAg-mediated toxic shock.
Publication types
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Amino Acid Sequence
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Animals
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Antigen Presentation / immunology*
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Antigens, Bacterial / immunology
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Antigens, Bacterial / metabolism*
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Antigens, Neoplasm / immunology
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Antigens, Neoplasm / metabolism
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Autoantigens / immunology
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Autoantigens / metabolism*
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Binding, Competitive / immunology
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Cell Membrane / immunology
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Cell Membrane / metabolism
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Enterotoxins / immunology
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Enterotoxins / metabolism*
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Female
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Histocompatibility Antigens Class II / biosynthesis
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Histocompatibility Antigens Class II / metabolism
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Histocompatibility Antigens Class II / physiology*
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Immune Tolerance*
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Male
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Mice
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Mice, Inbred A
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Mice, Inbred C57BL
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Mice, Knockout
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Molecular Sequence Data
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Protein Binding / immunology
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Staphylococcus aureus / immunology
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Superantigens / immunology
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Superantigens / metabolism*
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T-Lymphocytes / immunology
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T-Lymphocytes / metabolism
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T-Lymphocytes / microbiology
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Transfection
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Tumor Cells, Cultured
Substances
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Antigens, Bacterial
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Antigens, Neoplasm
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Autoantigens
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Enterotoxins
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H-2O antigen
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Histocompatibility Antigens Class II
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Superantigens
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enterotoxin B, staphylococcal