Abstract
The molecular adapter Fyb/Slap regulates signaling downstream of the T cell receptor (TCR), but whether it plays a positive or negative role is controversial. We demonstrate that Fyb/Slap-deficient T cells exhibit defective proliferation and cytokine production in response to TCR stimulation. Fyb/Slap is also required in vivo for T cell-dependent immune responses. Functionally, Fyb/Slap has no apparent role in the activation of known TCR signaling pathways, F-actin polymerization, or TCR clustering. Rather, Fyb/Slap regulates TCR-induced integrin clustering and adhesion. Thus, Fyb/Slap is the first molecular adapter to be identified that couples TCR stimulation to the avidity modulation of integrins governing T cell adhesion.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Actins / metabolism
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Adaptor Proteins, Signal Transducing*
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Animals
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Antigens, CD / metabolism
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Antigens, Differentiation, T-Lymphocyte / metabolism
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B-Lymphocytes / immunology
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CD3 Complex / metabolism
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Carrier Proteins / genetics
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Carrier Proteins / physiology*
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Cell Adhesion
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Cell Adhesion Molecules / metabolism
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Chimera
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Gene Targeting
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Humans
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Immunization
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Immunoglobulin G / biosynthesis
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Integrins / metabolism*
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Intercellular Adhesion Molecule-1 / metabolism
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Interferon-gamma / biosynthesis
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Interleukin-2 / biosynthesis
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Interleukin-2 / pharmacology
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Lectins, C-Type
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Lymphocyte Activation*
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Lymphocyte Function-Associated Antigen-1 / metabolism
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Mice
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Phosphoproteins / genetics
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Phosphoproteins / physiology*
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Receptors, Antigen, T-Cell / immunology
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Receptors, Antigen, T-Cell / metabolism
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Receptors, Interleukin-2 / metabolism
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Recombinant Proteins / metabolism
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Signal Transduction
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T-Lymphocytes / immunology
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T-Lymphocytes / metabolism
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T-Lymphocytes / physiology*
Substances
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Actins
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Adaptor Proteins, Signal Transducing
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Antigens, CD
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Antigens, Differentiation, T-Lymphocyte
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CD3 Complex
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CD69 antigen
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Carrier Proteins
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Cell Adhesion Molecules
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FYB1 protein, human
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Fyb protein, mouse
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ICAM2 protein, human
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Immunoglobulin G
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Integrins
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Interleukin-2
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Lectins, C-Type
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Lymphocyte Function-Associated Antigen-1
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Phosphoproteins
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Receptors, Antigen, T-Cell
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Receptors, Interleukin-2
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Recombinant Proteins
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Intercellular Adhesion Molecule-1
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Interferon-gamma