Abstract
The proinflammatory cytokine interleukin-1 (IL-1) transmits a signal via several critical cytoplasmic proteins such as MyD88, IRAKs and TRAF6. Recently, serine/threonine kinase TAK1 and TAK1 binding protein 1 and 2 (TAB1/2) have been identified as molecules involved in IL-1-induced TRAF6-mediated activation of AP-1 and NF-kappa B via mitogen-activated protein (MAP) kinases and I kappa B kinases, respectively. However, their physiological functions remain to be clarified. To elucidate their roles in vivo, we generated TAB2-deficient mice. The TAB2 deficiency was embryonic lethal due to liver degeneration and apoptosis. This phenotype was similar to that of NF-kappa B p65-, IKK beta-, and NEMO/IKK gamma-deficient mice. However, the IL-1-induced activation of NF-kappa B and MAP kinases was not impaired in TAB2-deficient embryonic fibroblasts. These findings demonstrate that TAB2 is essential for embryonic development through prevention of liver apoptosis but not for the IL-1 receptor-mediated signaling pathway.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Adaptor Proteins, Signal Transducing*
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Animals
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Apoptosis / genetics*
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Carrier Proteins / genetics
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Carrier Proteins / metabolism*
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Cells, Cultured
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Fetal Death / genetics
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Fibroblasts / drug effects
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Fibroblasts / physiology
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Genetic Engineering / methods
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Inflammation / metabolism
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Interleukin-1 / metabolism*
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Interleukin-1 / pharmacology
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Intracellular Signaling Peptides and Proteins*
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Liver / embryology*
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Liver / metabolism
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Liver / pathology*
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MAP Kinase Kinase Kinases / metabolism
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Mice
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Mice, Mutant Strains
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Mitogen-Activated Protein Kinases / drug effects
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Mitogen-Activated Protein Kinases / metabolism
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NF-kappa B / drug effects
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NF-kappa B / metabolism
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Phosphorylation
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Signal Transduction
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Tumor Necrosis Factor-alpha / metabolism
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Tumor Necrosis Factor-alpha / pharmacology
Substances
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Adaptor Proteins, Signal Transducing
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Carrier Proteins
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Interleukin-1
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Intracellular Signaling Peptides and Proteins
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NF-kappa B
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TAB1 protein, MAPKKK activator, vertebrate
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Tab2 protein, mouse
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Tumor Necrosis Factor-alpha
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Mitogen-Activated Protein Kinases
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MAP Kinase Kinase Kinases
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MAP kinase kinase kinase 7