Pdx1 expression in Irs2-deficient mouse beta-cells is regulated in a strain-dependent manner

J Biol Chem. 2003 Oct 31;278(44):43691-8. doi: 10.1074/jbc.M307004200. Epub 2003 Jul 16.

Abstract

We previously demonstrated that Irs2-/- mice develop diabetes due to beta-cell growth failure and insulin resistance; however, glucose-induced insulin secretion was increased in islets isolated from Irs2-/- mice. Pdx-1, a transcription factor important for maintenance of the beta-cell function, was recently reported to be severely reduced in Irs2-/- murine beta-cells. We report herein that Pdx-1 expression, including the amount of Pdx-1 localized in the nucleus, is not down-regulated in our Irs2-/- murine beta-cells with a C57BL/6 background. We have also demonstrated the expression of upstream genes of Pdx-1, such as HNF3beta and HNF1alpha, as well as its downstream genes, including insulin, Glut2, and Nkx6.1, to be well preserved. We have further demonstrated Pdx-1 expression to also be preserved in beta-cells of 30-week-old diabetic Irs2-/- mice. In addition, surprisingly, even in Irs2-/- mice on a high fat diet with markedly elevated blood glucose, exceeding 400 mg/dl, Pdx-1 expression was not reduced. Furthermore, we found Pdx-1 to be markedly decreased in certain severely diabetic Irs2-/- mice with a mixed C57BL/6J x 129Sv background. We conclude that 1) Pdx-1 expression in Irs2-/- mice is regulated in a strain-dependent manner, 2) Irs2-/- mice develop diabetes associated with beta-cell growth failure even when Pdx1 expression is preserved, and 3) Pdx-1 expression is preserved in severely hyperglycemic Irs2-/- mice with a C57BL/6 background on a high fat diet.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animal Feed
  • Animals
  • Blotting, Western
  • Down-Regulation
  • Female
  • Gene Expression Regulation*
  • Glucose / metabolism
  • Homeodomain Proteins / metabolism
  • Hypoglycemia / metabolism
  • Immunohistochemistry
  • Insulin / metabolism
  • Insulin Receptor Substrate Proteins
  • Insulin Secretion
  • Intracellular Signaling Peptides and Proteins
  • Islets of Langerhans / metabolism*
  • Male
  • Mice
  • Mice, Inbred C57BL
  • Mice, Transgenic
  • Phosphoproteins / genetics*
  • Phosphoproteins / physiology
  • RNA / metabolism
  • Reverse Transcriptase Polymerase Chain Reaction
  • Species Specificity
  • Time Factors
  • Trans-Activators / biosynthesis*
  • Trans-Activators / physiology*

Substances

  • Homeodomain Proteins
  • Insulin
  • Insulin Receptor Substrate Proteins
  • Intracellular Signaling Peptides and Proteins
  • Irs2 protein, mouse
  • Nkx6-1 protein, mouse
  • Phosphoproteins
  • Trans-Activators
  • pancreatic and duodenal homeobox 1 protein
  • RNA
  • Glucose