Abstract
Hydroxylation at an asparagine residue at the COOH-terminal activation domain of hypoxia-inducible factor (HIF)-1/2 alphas is essential for its inactivation under normoxic condition. To date, the mechanism by which HIF-alpha avoids the inhibitory effect of asparagine hydroxylase in renal cell carcinoma (RCC) in normoxia is undefined. We have shown herein that protein kinase C (PKC) zeta has an important role in HIF-alpha activation in RCC. By using dominant negative mutant and small interference RNA approaches, we have demonstrated that the association between HIF-alpha and p300 is modulated by PKCzeta. Moreover, a novel signaling pathway involving phosphatidylinositol 3'-kinase and PKCzeta has been shown to be responsible for the activation of HIF-alpha by inhibiting the mRNA expression of FIH-1 (factor inhibiting HIF-1) in RCC and thereby promoting the transcription of hypoxia-inducible genes such as vascular permeability factor/vascular endothelial growth factor.
Publication types
-
Research Support, Non-U.S. Gov't
-
Research Support, U.S. Gov't, P.H.S.
MeSH terms
-
Acetyltransferases / metabolism
-
Carcinoma, Renal Cell / enzymology*
-
Carcinoma, Renal Cell / pathology
-
Cell Cycle Proteins / metabolism
-
Cell Line, Tumor
-
DNA Primers
-
Gene Expression Regulation, Neoplastic
-
Histone Acetyltransferases
-
Humans
-
Hydroxylation
-
Hypoxia-Inducible Factor 1, alpha Subunit
-
Kidney Neoplasms / enzymology*
-
Kidney Neoplasms / pathology
-
Luciferases / genetics
-
Protein Kinase C / genetics*
-
RNA, Messenger / genetics
-
RNA, Small Interfering / genetics
-
Reverse Transcriptase Polymerase Chain Reaction
-
Transcription Factors / genetics
-
Transcription Factors / metabolism*
-
Transcription, Genetic
-
Transcriptional Activation
-
Transfection
-
p300-CBP Transcription Factors
Substances
-
Cell Cycle Proteins
-
DNA Primers
-
HIF1A protein, human
-
Hypoxia-Inducible Factor 1, alpha Subunit
-
RNA, Messenger
-
RNA, Small Interfering
-
Transcription Factors
-
Luciferases
-
Acetyltransferases
-
Histone Acetyltransferases
-
p300-CBP Transcription Factors
-
p300-CBP-associated factor
-
protein kinase C zeta
-
Protein Kinase C