Abstract
The oncoprotein HER-2/neu is a prosurvival factor, and its overexpression has been correlated with poor prognosis in patients with breast cancer. We report that HER-2 is a new substrate for caspase-8 and that tumor necrosis factor alpha (TNF-alpha) stimulation leads to an early caspase-8-dependent HER-2 cleavage in MCF7 A/Z breast adenocarcinoma cells defective for nuclear factor kappaB (NFkappaB) activation. We show that the antiapoptotic transcription factor NFkappaB counteracts this cleavage through induction of the caspase-8 inhibitor c-FLIP. Our results also demonstrate that this HER-2 cleavage contributes to the TNF-alpha-induced apoptosis pathway because ectopic expression of an uncleavable HER-2 protects NFkappaB-defective cells against TNF-alpha-mediated cell death. Therefore, we propose an original model in which NFkappaB exerts a new antiapoptotic function by counteracting TNF-alpha-triggered cleavage of the HER-2 survival factor.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Adenocarcinoma / enzymology
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Adenocarcinoma / metabolism*
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Adenocarcinoma / pathology
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Anti-Infective Agents / pharmacology
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Apoptosis / physiology
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Binding Sites
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Breast Neoplasms / enzymology
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Breast Neoplasms / metabolism*
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Breast Neoplasms / pathology
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CASP8 and FADD-Like Apoptosis Regulating Protein
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Carrier Proteins / metabolism*
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Caspase 8
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Caspase Inhibitors
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Caspases / metabolism*
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Cell Line, Tumor
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Cycloheximide / pharmacology
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Enzyme Activation
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Humans
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Intracellular Signaling Peptides and Proteins*
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NF-kappa B / antagonists & inhibitors
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NF-kappa B / metabolism*
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Nitriles
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Receptor, ErbB-2 / biosynthesis
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Receptor, ErbB-2 / genetics
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Receptor, ErbB-2 / metabolism*
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Stimulation, Chemical
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Sulfones
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Transcriptional Activation
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Tumor Necrosis Factor-alpha / pharmacology*
Substances
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Anti-Infective Agents
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BAY 11-7085
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CASP8 and FADD-Like Apoptosis Regulating Protein
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CFLAR protein, human
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Carrier Proteins
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Caspase Inhibitors
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Intracellular Signaling Peptides and Proteins
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NF-kappa B
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Nitriles
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Sulfones
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Tumor Necrosis Factor-alpha
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Cycloheximide
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Receptor, ErbB-2
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CASP8 protein, human
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Caspase 8
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Caspases