Abstract
Neuregulin-1 (NRG-1) growth and differentiation factors and their erbB receptors are hypothesized to promote embryonic hippocampal neuron differentiation via as yet unknown mechanisms. We have found that NRG-1beta increases the outgrowth of primary neurites, neuronal area, total neurite length, and neuritic branching in E18 hippocampal neurons. NRG-1beta effects on neurite extension and arborization are similar to, but not additive with, those of brain-derived neurotrophic factor and reflect direct NRG-1 action on hippocampal neurons as these cells express the NRG-1 receptors erbB2 and erbB4, the erbB-specific inhibitor PD158780 decreases NRG-1beta induced neurite outgrowth, and NRG-1beta stimulation induces p42/44 ERK phosphorylation. Pharmacological inhibition of p42/44 ERK and protein kinase C (PKC), but not PI3K or p38 MAP kinase, inhibits NRG-1beta-induced neurite extension and elaboration. We conclude that NRG-1beta stimulates hippocampal neurite extension and arborization via a signaling pathway that involves erbB membrane tyrosine kinases (erbB2 and/or erbB4), p42/44 ERK, and PKC.
Publication types
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Animals
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Brain-Derived Neurotrophic Factor / metabolism
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Brain-Derived Neurotrophic Factor / pharmacology
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Cell Differentiation / drug effects
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Cell Differentiation / physiology*
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Cells, Cultured
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Enzyme Inhibitors / pharmacology
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ErbB Receptors / metabolism
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Hippocampus / cytology
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Hippocampus / embryology*
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Hippocampus / metabolism
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Mitogen-Activated Protein Kinase 1 / antagonists & inhibitors
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Mitogen-Activated Protein Kinase 1 / metabolism
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Nerve Tissue Proteins / pharmacology
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Nerve Tissue Proteins / physiology*
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Neuregulin-1
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Neurites / drug effects
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Neurites / metabolism*
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Neurites / ultrastructure
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Protein Isoforms / pharmacology
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Protein Isoforms / physiology
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Protein Kinase C / antagonists & inhibitors
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Protein Kinase C / metabolism
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Rats
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Receptor, ErbB-2 / metabolism
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Receptor, ErbB-4
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Signal Transduction / physiology
Substances
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Brain-Derived Neurotrophic Factor
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Enzyme Inhibitors
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Nerve Tissue Proteins
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Neuregulin-1
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Nrg1 protein, rat
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Protein Isoforms
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ErbB Receptors
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Erbb4 protein, rat
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Receptor, ErbB-2
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Receptor, ErbB-4
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Protein Kinase C
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Mitogen-Activated Protein Kinase 1