Abstract
Interphotoreceptor retinoid binding protein (IRBP)-induced experimental autoimmune uveoretinitis (EAU) is a CD4+ T cell-mediated autoimmune disease. Development of EAU is inhibited by treatment with an agonistic anti-4-1BB mAb. Even established EAU was alleviated by anti-4-1BB mAb. However, inhibition of 4-1BB/4-1BB ligand (4-1BBL) interaction does not suppress the development of EAU. It appears that cross-linking of 4-1BB evokes an active antigen-specific suppression mechanism rather than merely blocking 4-1BB/4-1BBL interaction. We found that administration of anti-4-1BB mAb induced massive clonal expansion of CD11c+CD8+ T cells that produced IFN-gamma, resulting in accumulation of a high level of indoleamine 2,3-dioxygenase (IDO) in CD11c+ dendritic cells. 4-1BB-mediated suppression of EAU was reversed by the pharmacological IDO inhibitor, 1-methyl-tryptophan (1-MT). These studies demonstrate that suppression of EAU results from antigen-driven, 4-1BB-mediated expansion of novel CD11c+CD8+ T cells that suppress antigen-specific CD4+ T cells via an IDO-dependent mechanism.
Publication types
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Research Support, N.I.H., Extramural
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Research Support, Non-U.S. Gov't
MeSH terms
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4-1BB Ligand
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Animals
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Autoimmune Diseases / enzymology
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Autoimmune Diseases / genetics
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Autoimmune Diseases / immunology
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Autoimmune Diseases / metabolism*
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CD11 Antigens / metabolism
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CD8-Positive T-Lymphocytes / cytology
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CD8-Positive T-Lymphocytes / metabolism
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Cell Proliferation
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Disease Models, Animal
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Enzyme Inhibitors / pharmacology
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Indoleamine-Pyrrole 2,3,-Dioxygenase / antagonists & inhibitors
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Indoleamine-Pyrrole 2,3,-Dioxygenase / metabolism*
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Inflammation / immunology
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Inflammation / metabolism
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Inflammation / pathology
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Interferon-gamma / biosynthesis
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Mice
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Mice, Inbred C57BL
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Mice, Knockout
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Nitric Oxide Synthase Type II / antagonists & inhibitors
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Nitric Oxide Synthase Type II / metabolism
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Retina / metabolism*
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Retina / pathology*
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Tumor Necrosis Factors / deficiency
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Tumor Necrosis Factors / genetics
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Tumor Necrosis Factors / immunology
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Tumor Necrosis Factors / metabolism*
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Uvea / metabolism*
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Uvea / pathology*
Substances
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4-1BB Ligand
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CD11 Antigens
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Enzyme Inhibitors
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Indoleamine-Pyrrole 2,3,-Dioxygenase
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Tnfsf9 protein, mouse
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Tumor Necrosis Factors
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Interferon-gamma
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Nitric Oxide Synthase Type II