Loss of M5 muscarinic acetylcholine receptors leads to cerebrovascular and neuronal abnormalities and cognitive deficits in mice

Neurobiol Dis. 2006 Nov;24(2):334-44. doi: 10.1016/j.nbd.2006.07.010. Epub 2006 Sep 7.

Abstract

The M5 muscarinic acetylcholine receptor (M5R) has been shown to play a crucial role in mediating acetylcholine-dependent dilation of cerebral blood vessels. We show that male M5R-/- mice displayed constitutive constriction of cerebral arteries using magnetic resonance angiography in vivo. Male M5R-/- mice exhibited a significantly reduced cerebral blood flow (CBF) in the cerebral cortex, hippocampus, basal ganglia, and thalamus. Cortical and hippocampal pyramidal neurons from M5R-/- mice showed neuronal atrophy. Hippocampus-dependent spatial and nonspatial memory was also impaired in M5R-/- mice. In M5R-/- mice, CA3 pyramidal cells displayed a significantly attenuated frequency of the spontaneous postsynaptic current and long-term potentiation was significantly impaired at the mossy fiber-CA3 synapse. Our findings suggest that impaired M5R signaling may play a role in the pathophysiology of cerebrovascular deficits. The M5 receptor may represent an attractive novel therapeutic target to ameliorate memory deficits caused by impaired cerebrovascular function.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Acetylcholine / metabolism
  • Animals
  • Brain / blood supply
  • Brain / metabolism*
  • Brain / physiopathology
  • Cerebral Arteries / metabolism*
  • Cerebral Arteries / pathology
  • Cerebral Arteries / physiopathology
  • Cerebral Cortex / metabolism
  • Cerebral Cortex / pathology
  • Cerebral Cortex / physiopathology
  • Cerebrovascular Circulation / genetics
  • Cerebrovascular Disorders / genetics
  • Cerebrovascular Disorders / metabolism*
  • Cerebrovascular Disorders / physiopathology
  • Cognition Disorders / genetics
  • Cognition Disorders / metabolism*
  • Cognition Disorders / physiopathology
  • Disease Models, Animal
  • Hippocampus / metabolism
  • Hippocampus / pathology
  • Hippocampus / physiopathology
  • Long-Term Potentiation / genetics
  • Magnetic Resonance Angiography
  • Male
  • Memory Disorders / genetics
  • Memory Disorders / metabolism
  • Memory Disorders / physiopathology
  • Mice
  • Mice, Inbred C57BL
  • Mice, Knockout
  • Nerve Degeneration / genetics
  • Nerve Degeneration / metabolism
  • Nerve Degeneration / pathology
  • Neurodegenerative Diseases / genetics
  • Neurodegenerative Diseases / metabolism*
  • Neurodegenerative Diseases / physiopathology
  • Pyramidal Cells / metabolism
  • Pyramidal Cells / pathology
  • Receptor, Muscarinic M5 / genetics*
  • Synaptic Transmission / genetics
  • Vasoconstriction / genetics

Substances

  • Receptor, Muscarinic M5
  • Acetylcholine