Allergenicity and antigenicity of wild-type and mutant, monomeric, and dimeric carrot major allergen Dau c 1: destruction of conformation, not oligomerization, is the roadmap to save allergen vaccines

J Allergy Clin Immunol. 2007 Apr;119(4):944-51. doi: 10.1016/j.jaci.2006.11.699. Epub 2007 Feb 9.

Abstract

Background: Carrot allergy is caused by primary sensitization to birch pollen. Continuous carrot exposure results in additional Dau c 1-specific allergic responses. Thus, immunotherapy with birch pollen may not improve the food allergy.

Objective: Evaluation of mutation and oligomerization of the major carrot allergen, Dau c 1, in regard to alteration of antibody binding capacities, structure, and the ability to induce blocking IgG antibodies.

Methods: Measurement of IgE reactivities to monomers, dimers of wild-type and mutant Dau c 1.0104 and Dau c 1.0201, and Dau c 1.0104 trimer, their ability to induce blocking antibodies in mice, and their allergenic potency by histamine release.

Results: The reactivity of human IgE to the mutant dimer was reduced on average by 81%. Sera of immunized Balb/c mice showed specific IgG similar to the human IgE antibody response; Dau c 1.01 was more antigenic than Dau c 1.02. Both wild-type and mutant Dau c 1 variants induced cross-reacting IgG, which blocked binding of human IgE. The mutants were more antigenic than the wild-type forms, and the dimers induced higher IgG responses in mice than the monomers. The results of the histamine release experiments corroborated the findings of the antibody binding studies.

Conclusion: Destruction of native conformation rather than oligomerization is the appropriate strategy to reduce the allergenicity of Bet v 1-homologous food allergens.

Clinical implications: The dimer composed of mutants of Dau c 1.0104 and Dau c 1.0201 is a promising candidate vaccine for treatment of carrot allergy because of its high immunogenicity and drastically reduced allergenicity.

MeSH terms

  • Allergens / administration & dosage
  • Allergens / chemistry
  • Allergens / genetics*
  • Allergens / immunology*
  • Animals
  • Antigens, Plant / administration & dosage
  • Antigens, Plant / chemistry
  • Antigens, Plant / genetics*
  • Antigens, Plant / immunology*
  • Dimerization
  • Dose-Response Relationship, Immunologic
  • Female
  • Humans
  • Immunoglobulin E / biosynthesis
  • Mice
  • Mice, Inbred BALB C
  • Plant Proteins / administration & dosage
  • Plant Proteins / chemistry
  • Plant Proteins / genetics*
  • Plant Proteins / immunology*
  • Protein Conformation
  • Protein Isoforms / administration & dosage
  • Protein Isoforms / chemistry
  • Protein Isoforms / genetics
  • Protein Isoforms / immunology
  • Vaccines, DNA / administration & dosage*
  • Vaccines, DNA / chemical synthesis
  • Vaccines, DNA / immunology*

Substances

  • Allergens
  • Antigens, Plant
  • Dau c 1 protein, Daucus carota
  • Plant Proteins
  • Protein Isoforms
  • Vaccines, DNA
  • Immunoglobulin E