Oxyntomodulin differentially affects glucagon-like peptide-1 receptor beta-arrestin recruitment and signaling through Galpha(s)

J Pharmacol Exp Ther. 2007 Jul;322(1):148-54. doi: 10.1124/jpet.107.120006. Epub 2007 Mar 29.

Abstract

The glucagon-like peptide (GLP)-1 receptor is a promising target for the treatment of type 2 diabetes and obesity, and there is great interest in characterizing the pharmacology of the GLP-1 receptor and its ligands. In the present report, we have applied bioluminescence resonance energy transfer assays to measure agonist-induced recruitment of betaarrestins and G-protein-coupled receptor kinase (GRK) 2 to the GLP-1 receptor in addition to traditional measurements of second messenger generation. The peptide hormone oxyntomodulin is described in the literature as a full agonist on the glucagon and GLP-1 receptors. Surprisingly, despite being full agonists in GLP-1 receptor-mediated cAMP accumulation, oxyntomodulin and glucagon were observed to be partial agonists in recruiting betaarrestins and GRK2 to the GLP-1 receptor. We suggest that oxyntomodulin and glucagon are biased ligands on the GLP-1 receptor.

MeSH terms

  • Animals
  • Arrestins / metabolism*
  • COS Cells
  • Chlorocebus aethiops
  • Cyclic AMP / metabolism
  • G-Protein-Coupled Receptor Kinase 2
  • G-Protein-Coupled Receptor Kinase 5
  • GTP-Binding Protein alpha Subunits, Gs / physiology*
  • Glucagon / pharmacology
  • Glucagon-Like Peptide 1 / pharmacology
  • Glucagon-Like Peptide-1 Receptor
  • Humans
  • Luminescent Measurements
  • Oxyntomodulin / pharmacology*
  • Protein Serine-Threonine Kinases / physiology
  • Receptors, Glucagon / drug effects*
  • Receptors, Glucagon / physiology
  • Signal Transduction / physiology*
  • beta-Adrenergic Receptor Kinases / physiology
  • beta-Arrestins

Substances

  • Arrestins
  • GLP1R protein, human
  • Glucagon-Like Peptide-1 Receptor
  • Oxyntomodulin
  • Receptors, Glucagon
  • beta-Arrestins
  • Glucagon-Like Peptide 1
  • Glucagon
  • Cyclic AMP
  • Protein Serine-Threonine Kinases
  • GRK2 protein, human
  • beta-Adrenergic Receptor Kinases
  • G-Protein-Coupled Receptor Kinase 2
  • G-Protein-Coupled Receptor Kinase 5
  • GRK5 protein, human
  • GTP-Binding Protein alpha Subunits, Gs