The bacterium, nontypeable Haemophilus influenzae, enhances host antiviral response by inducing Toll-like receptor 7 expression: evidence for negative regulation of host anti-viral response by CYLD

FEBS J. 2007 Jul;274(14):3655-3668. doi: 10.1111/j.1742-4658.2007.05899.x. Epub 2007 Jul 3.

Abstract

The incidence of mixed viral/bacterial infections has increased recently because of the dramatic increase in antibiotic-resistant strains, the emergence of new pathogens, and the resurgence of old ones. Despite the relatively well-known role of viruses in enhancing bacterial infections, the impact of bacterial infections on viral infections remains unknown. In this study, we provide direct evidence that nontypeable Haemophilus influenzae (NTHi), a major respiratory bacterial pathogen, augments the host antiviral response by up-regulating epithelial Toll-like receptor 7 (TLR7) expression in vitro and in vivo. Moreover, NTHi induces TLR7 expression via a TLR2-MyD88-IRAK-TRAF6-IKK-NF-kappaB-dependent signaling pathway. Interestingly, CYLD, a novel deubiquitinase, acts as a negative regulator of TLR7 induction by NTHi. Our study thus provides new insights into a novel role for bacterial infection in enhancing host antiviral response and further identifies CYLD for the first time as a critical negative regulator of host antiviral response.

Publication types

  • Research Support, N.I.H., Extramural

MeSH terms

  • Animals
  • Cell Line
  • Cysteine Endopeptidases / deficiency
  • Cysteine Endopeptidases / genetics
  • Cysteine Endopeptidases / metabolism*
  • Cytokines / biosynthesis
  • Deubiquitinating Enzyme CYLD
  • Down-Regulation*
  • Haemophilus influenzae / classification
  • Haemophilus influenzae / physiology*
  • Humans
  • Lung Diseases / genetics
  • Lung Diseases / metabolism
  • Lung Diseases / microbiology
  • Lung Diseases / virology
  • Membrane Glycoproteins / genetics
  • Membrane Glycoproteins / metabolism*
  • Mice
  • Signal Transduction
  • Toll-Like Receptor 2 / deficiency
  • Toll-Like Receptor 2 / genetics
  • Toll-Like Receptor 2 / metabolism
  • Toll-Like Receptor 7 / genetics
  • Toll-Like Receptor 7 / metabolism*
  • Tumor Suppressor Proteins / genetics
  • Tumor Suppressor Proteins / metabolism*
  • Up-Regulation*

Substances

  • Cytokines
  • Membrane Glycoproteins
  • TLR2 protein, human
  • TLR7 protein, human
  • Tlr2 protein, mouse
  • Tlr7 protein, mouse
  • Toll-Like Receptor 2
  • Toll-Like Receptor 7
  • Tumor Suppressor Proteins
  • CYLD protein, human
  • CYLD protein, mouse
  • Deubiquitinating Enzyme CYLD
  • Cysteine Endopeptidases