The zinc-binding motif of human RECQ5beta suppresses the intrinsic strand-annealing activity of its DExH helicase domain and is essential for the helicase activity of the enzyme

Biochem J. 2008 Jun 15;412(3):425-33. doi: 10.1042/BJ20071150.

Abstract

RecQ family helicases, functioning as caretakers of genomic integrity, contain a zinc-binding motif which is highly conserved among these helicases, but does not have a substantial structural similarity with any other known zinc-finger folds. In the present study, we show that a truncated variant of the human RECQ5beta helicase comprised of the conserved helicase domain only, a splice variant named RECQ5alpha, possesses neither ATPase nor DNA-unwinding activities, but surprisingly displays a strong strand-annealing activity. In contrast, fragments of RECQ5beta including the intact zinc-binding motif, which is located immediately downstream of the helicase domain, exhibit much reduced strand-annealing activity but are proficient in DNA unwinding. Quantitative measurements indicate that the regulatory role of the zinc-binding motif is achieved by enhancing the DNA-binding affinity of the enzyme. The novel intramolecular modulation of RECQ5beta catalytic activity mediated by the zinc-binding motif may represent a universal regulation mode for all RecQ family helicases.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adenosine Triphosphatases / metabolism
  • Adenosine Triphosphate / metabolism
  • Amino Acid Motifs
  • Amino Acid Sequence
  • DNA, Single-Stranded / metabolism
  • Genetic Variation
  • Humans
  • Models, Biological
  • Models, Molecular
  • Molecular Sequence Data
  • RecQ Helicases / chemistry*
  • RecQ Helicases / genetics
  • RecQ Helicases / metabolism*
  • Zinc Fingers*

Substances

  • DNA, Single-Stranded
  • RECQL5 protein, human
  • Adenosine Triphosphate
  • Adenosine Triphosphatases
  • RecQ Helicases