Reduction in hypophyseal growth hormone and prolactin expression due to deficiency in ghrelin receptor signaling is associated with Pit-1 suppression: relevance to the immune system

Brain Behav Immun. 2008 Nov;22(8):1138-45. doi: 10.1016/j.bbi.2008.06.003. Epub 2008 Jun 17.

Abstract

In mice and in rats, reduced levels of the growth hormone secretagogue receptor (GHS-R1a) results in reduced body weight and lower levels of serum insulin-like growth factor I (IGF-I). However, the mechanism leading to these impairments has not been elucidated. Studies in primary cultures of pituitary cells from very young mice have shown that GHS-R1a agonists, including ghrelin, increase expression of the pituitary-specific transcription factor (Pit-1) that is critical for differentiation of pituitary cells into somatotrophs, lactotrophs, and thyrotrophs. Hence, we hypothesized that ablation of Ghsr would reduce Pit-1 expression and as a consequence reduce growth hormone (GH) production explaining the lower body weight of Ghsr-/- mice. Here, we now show that Pit-1 mRNA levels are significantly lower in the pituitary gland of Ghsr-/- mice compared to wild-type littermates and also with advancing age. This Pit-1 loss is associated with reduced GH mRNA and fewer GH producing cells. To determine whether reduced GH is caused by reduced expression of Pit-1 in Ghsr-/- mice, we also measured prolactin (PRL) expression in the pituitary gland and in the circulation. PRL mRNA was significantly reduced in Ghsr-/- mice compared to wild-type littermates and fewer cells expressed PRL. The reduction in expression of both GH and PRL is consistent with a Pit-1 regulated pathway and demonstrates that the GHS-R has an important role in the pituitary gland as a modulator of Pit-1 expression and provides a possible mechanism to explain the lower plasma IGF-1 and modestly reduced body weight exhibited by Ghsr-/- mice. We also believe that lower systemic and lymphoid hormone expression may also account, in part, for the enhanced thymic involution and reduced thymic output in Ghsr-/- mice.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, N.I.H., Intramural

MeSH terms

  • Age Factors
  • Analysis of Variance
  • Animals
  • Enzyme-Linked Immunosorbent Assay
  • Female
  • Growth Hormone / metabolism*
  • Immunohistochemistry
  • Insulin-Like Growth Factor I / metabolism
  • Male
  • Mice
  • Mice, Transgenic
  • Organ Size
  • Pituitary Gland / growth & development*
  • Pituitary Gland / metabolism*
  • Prolactin / blood
  • Prolactin / metabolism*
  • RNA, Messenger / metabolism
  • Receptors, Ghrelin / genetics*
  • Receptors, Ghrelin / metabolism
  • Reverse Transcriptase Polymerase Chain Reaction
  • Transcription Factor Pit-1 / metabolism*

Substances

  • RNA, Messenger
  • Receptors, Ghrelin
  • Transcription Factor Pit-1
  • Insulin-Like Growth Factor I
  • Prolactin
  • Growth Hormone