Abstract
Nuclear orphan receptor Nur77 has important roles in many biological processes. However, a physiological ligand for Nur77 has not been identified. Here, we report that the octaketide cytosporone B (Csn-B) is a naturally occurring agonist for Nur77. Csn-B specifically binds to the ligand-binding domain of Nur77 and stimulates Nur77-dependent transactivational activity towards target genes including Nr4a1 (Nur77) itself, which contains multiple consensus response elements allowing positive autoregulation in a Csn-B-dependent manner. Csn-B also elevates blood glucose levels in fasting C57 mice, an effect that is accompanied by induction of multiple genes involved in gluconeogenesis. These biological effects were not observed in Nur77-null (Nr4a1-/-) mice, which indicates that Csn-B regulates gluconeogenesis through Nur77. Moreover, Csn-B induced apoptosis and retarded xenograft tumor growth by inducing Nur77 expression, translocating Nur77 to mitochondria to cause cytochrome c release. Thus, Csn-B may represent a promising therapeutic drug for cancers and hypoglycemia, and it may also be useful as a reagent to increase understanding of Nur77 biological function.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Antineoplastic Agents* / isolation & purification
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Antineoplastic Agents* / pharmacology
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Antineoplastic Agents* / therapeutic use
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Apoptosis / drug effects
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Ascomycota / chemistry
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Blood Glucose / metabolism
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Cell Line, Tumor
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Cell Proliferation / drug effects
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DNA-Binding Proteins / agonists*
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DNA-Binding Proteins / biosynthesis
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DNA-Binding Proteins / genetics
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Gluconeogenesis / drug effects*
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Gluconeogenesis / genetics
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Humans
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Ligands
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Mice
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Mice, Inbred BALB C
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Mice, Inbred C57BL
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Mice, Nude
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Models, Molecular
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Neoplasm Transplantation
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Neoplasms / drug therapy*
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Neoplasms / metabolism
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Neoplasms / pathology
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Nuclear Receptor Subfamily 4, Group A, Member 1
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Phenylacetates* / isolation & purification
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Phenylacetates* / pharmacology
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Phenylacetates* / therapeutic use
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Protein Binding
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Protein Transport
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Receptors, Steroid / agonists*
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Receptors, Steroid / biosynthesis
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Receptors, Steroid / genetics
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Transcriptional Activation
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Transfection
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Xenograft Model Antitumor Assays
Substances
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Antineoplastic Agents
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Blood Glucose
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DNA-Binding Proteins
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Ligands
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NR4A1 protein, human
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Nr4a1 protein, mouse
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Nuclear Receptor Subfamily 4, Group A, Member 1
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Phenylacetates
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Receptors, Steroid
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cytosporone B