A sperm GPI-anchored protein elicits sperm-cumulus cross-talk leading to the acrosome reaction

Cell Mol Life Sci. 2009 Mar;66(5):900-8. doi: 10.1007/s00018-009-8482-2.

Abstract

The acrosome reaction has long been thought to be induced by the zona pellucida. Here we report the identification and function of a novel human sperm glycosylphosphatidylinositol (GPI)-anchored membrane protein, NYD-SP8. The release of the protein during sperm-egg interaction and its binding to the cumulus, the first layer of egg investment, elicits cross-talk between the gametes and produces calcium dependant release of progesterone, which lead to the acrosome reaction. An in vivo mouse model of NYD-SP8 immunization is also established showing a reduced fertility rate. Thus, contrary to accepted dogma, our study demonstrates for the first time that, prior to reaching the zona pellucida, sperm may release a surface protein that acts on the cumulus cells leading to the acrosome reaction, which may be important for determining the outcome of fertilization.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Acrosome Reaction / physiology*
  • Amino Acid Sequence
  • Animals
  • Antibodies, Monoclonal / genetics
  • Antibodies, Monoclonal / metabolism
  • Antigens, Surface / genetics
  • Antigens, Surface / metabolism
  • Calcium / metabolism
  • Cell Communication / physiology*
  • Cumulus Cells / cytology
  • Cumulus Cells / metabolism*
  • Female
  • Fertility
  • Fertilization
  • GPI-Linked Proteins
  • Glycosylphosphatidylinositols / genetics
  • Glycosylphosphatidylinositols / metabolism*
  • Humans
  • Male
  • Membrane Proteins / genetics
  • Membrane Proteins / metabolism*
  • Mice
  • Molecular Sequence Data
  • Oocytes / cytology
  • Oocytes / metabolism
  • Progesterone / metabolism
  • Sequence Alignment
  • Spermatozoa / cytology
  • Spermatozoa / metabolism*
  • Tissue Distribution

Substances

  • Antibodies, Monoclonal
  • Antigens, Surface
  • GPI-Linked Proteins
  • Glycosylphosphatidylinositols
  • Membrane Proteins
  • TEX101 protein, human
  • TEX101 protein, mouse
  • Progesterone
  • Calcium