Abstract
Although rhein has been shown to induce apoptosis in several cancer cell lines, the mechanism of action of rhein-induced cell cycle arrest and apoptosis at the molecular level is not well known. In this study, the mechanism of rhein action on A-549 human lung cancer cells was investigated. Rhein induced G0/G1 arrest through inhibition of cyclin D3, Cdk4 and Cdk6. The efficacious induction of apoptosis was observed at 50 microM for 12 h and up to 72 h as examined by a flow cytometric method. Flow cytometric analysis demonstrated that rhein increased the levels of GADD153 and GRP78, both hallmarks of endoplasmic reticulum stress, promoted ROS and Ca2+ production, induced the loss of mitochondrial membrane potential (delta psi(m)), promoted cytochrome c release from mitochondria, promoted capase-3 activation and led to apoptosis. Rhein also increased the levels of p53, p21 and Bax but reduced the level of Bcl-2. The Ca2+ chelator BAPTA was added to the cells before rhein treatment, thus blocking the Ca2+ production and inhibiting rhein-induced apoptosis in A-549 cells. Our data demonstrate that rhein induces apoptosis in A-549 cells via a Ca2+ -dependent mitochondrial pathway.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Anthraquinones / pharmacology*
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Apoptosis / drug effects*
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Apoptosis / physiology
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Blotting, Western
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Calcium / metabolism*
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Caspase Inhibitors
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Cell Line, Tumor
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Cyclin D3
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Cyclin-Dependent Kinase 4 / antagonists & inhibitors
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Cyclin-Dependent Kinase 6 / antagonists & inhibitors
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Cyclins / antagonists & inhibitors
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Egtazic Acid / analogs & derivatives
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Egtazic Acid / pharmacology
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Endoplasmic Reticulum / drug effects*
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Endoplasmic Reticulum / metabolism
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Endoplasmic Reticulum Chaperone BiP
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Enzyme Inhibitors / pharmacology
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Flow Cytometry
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G1 Phase / drug effects
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Humans
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Lung Neoplasms / drug therapy*
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Lung Neoplasms / metabolism
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Lung Neoplasms / pathology
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Membrane Potential, Mitochondrial / drug effects
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Microscopy, Confocal
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Oligopeptides / pharmacology
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Reactive Oxygen Species / metabolism
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Resting Phase, Cell Cycle / drug effects
Substances
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Anthraquinones
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CCND3 protein, human
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Caspase Inhibitors
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Cyclin D3
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Cyclins
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Endoplasmic Reticulum Chaperone BiP
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Enzyme Inhibitors
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HSPA5 protein, human
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Oligopeptides
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Reactive Oxygen Species
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benzoylcarbonyl-aspartyl-glutamyl-valyl-aspartyl-fluoromethyl ketone
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Egtazic Acid
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CDK4 protein, human
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CDK6 protein, human
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Cyclin-Dependent Kinase 4
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Cyclin-Dependent Kinase 6
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1,2-bis(2-aminophenoxy)ethane-N,N,N',N'-tetraacetic acid
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Calcium
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rhein