Origin and evolution of the vertebrate leukocyte receptors: the lesson from tunicates

Immunogenetics. 2009 Jun;61(6):463-81. doi: 10.1007/s00251-009-0373-z. Epub 2009 Apr 30.

Abstract

Two selected receptor genes of the immunoglobulin superfamily (IgSF), one CTX/JAM family member, and one poliovirus receptor-like nectin that have features of adhesion molecules can be expressed by Ciona hemocytes, the effectors of immunity. They can also be expressed in the nervous system (CTX/JAM) and in the ovary (nectin). The genes encoding these receptors are located among one set of genes, spread over Ciona chromosomes 4 and 10, and containing other IgSF members homologous to those encoded by genes present in a tetrad of human (1, 3 + X, 11, 21 + 19q) or bird chromosomes (1, 4, 24, 31) that include the leukocyte receptor complex. It is proposed that this tetrad is due to the two rounds of duplication that affected a single prevertebrate ancestral region containing a primordial leukocyte receptor complex involved in immunity and other developmental regulatory functions.

MeSH terms

  • Amino Acid Sequence
  • Animals
  • Cell Adhesion Molecules / genetics
  • Chickens / genetics
  • Chromosome Mapping
  • Ciona intestinalis / embryology
  • Ciona intestinalis / genetics
  • Ciona intestinalis / growth & development
  • Evolution, Molecular*
  • Female
  • Gene Expression Regulation, Developmental
  • Hemocytes / metabolism
  • Histocompatibility Antigens Class I / genetics
  • In Situ Hybridization
  • Junctional Adhesion Molecules
  • Leukocytes / immunology
  • Leukocytes / metabolism
  • Male
  • Molecular Sequence Data
  • Nectins
  • Phylogeny*
  • Receptors, Immunologic / classification
  • Receptors, Immunologic / genetics*
  • Reverse Transcriptase Polymerase Chain Reaction
  • Sequence Homology, Amino Acid
  • Synteny
  • Urochordata / embryology
  • Urochordata / genetics
  • Urochordata / growth & development
  • Vertebrates / classification
  • Vertebrates / genetics*
  • Vertebrates / immunology

Substances

  • Cell Adhesion Molecules
  • Histocompatibility Antigens Class I
  • Junctional Adhesion Molecules
  • Nectins
  • Receptors, Immunologic