Signaling of c-kit in dendritic cells influences adaptive immunity

Ann N Y Acad Sci. 2010 Jan:1183:104-22. doi: 10.1111/j.1749-6632.2009.05122.x.

Abstract

The binding of the receptor tyrosine kinase, c-kit, to its ligand, stem cell factor (SCF), mediates numerous biological functions. Important roles for c-kit in hematopoiesis, melanogenesis, erythropoiesis, spermatogenesis, and carcinogenesis are well documented. Similarly, activation of granulocytes, mast cells, and of eosinophils in particular, by c-kit ligation has long been known to result in degranulation with concomitant release of pro-inflammatory mediators, including cytokines. However, recent work from a number of laboratories, including our own, highlights previously unappreciated functions for c-kit in immunologic processes. These novel findings strongly suggest that signaling through the c-kit-SCF axis could have a significant impact on the pathogenesis of diseases associated with an immunologic component. In our own studies, c-kit upregulation on dendritic cells via T helper (Th)2- and Th17-inducing stimuli led to c-kit activation and immune skewing toward these T helper subsets and away from Th1 responses. Others have shown that dendritic cell treatment with inhibitors of c-kit activation, such as imatinib mesylate (Gleevec), favored breaking of T-cell tolerance, skewing of responses toward production of Th1 cytokines, and activation of natural killer cells. These data all indicate that deeper understanding of, and ability to control, the c-kit-SCF axis could lead to improved treatment modalities aimed at redirecting unwanted and/or deleterious immune responses in a wide variety of conditions.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't
  • Review

MeSH terms

  • Adaptive Immunity / genetics
  • Adaptive Immunity / physiology*
  • Animals
  • Cell Membrane / metabolism
  • Dendritic Cells / immunology*
  • Dendritic Cells / metabolism
  • Hematopoietic Stem Cells / metabolism
  • Hematopoietic Stem Cells / physiology
  • Humans
  • Models, Biological
  • Protein Isoforms / genetics
  • Protein Isoforms / metabolism
  • Protein Isoforms / physiology
  • Proto-Oncogene Proteins c-kit / genetics
  • Proto-Oncogene Proteins c-kit / metabolism
  • Proto-Oncogene Proteins c-kit / physiology*
  • Receptors, Notch / metabolism
  • Receptors, Notch / physiology
  • Signal Transduction / genetics
  • Signal Transduction / immunology
  • Stem Cell Factor / genetics
  • Stem Cell Factor / metabolism
  • Stem Cell Factor / physiology

Substances

  • Protein Isoforms
  • Receptors, Notch
  • Stem Cell Factor
  • Proto-Oncogene Proteins c-kit