Abstract
The Salvador (Sav)/Warts (Wts)/Hippo (Hpo) (SWH) network controls tissue growth by inhibiting cell proliferation and promoting apoptosis. The core of the pathway consists of a MST and LATS family kinase cascade that ultimately phosphorylates and inactivates the YAP/Yorkie (Yki) transcription coactivator. The FERM domain proteins Merlin (Mer) and Expanded (Ex) represent one mode of upstream regulation controlling pathway activity. Here, we identify Kibra as a member of the SWH network. Kibra, which colocalizes and associates with Mer and Ex, also promotes the Mer/Ex association. Furthermore, the Kibra/Mer association is conserved in human cells. Finally, Kibra complexes with Wts and kibra depletion in tissue culture cells induces a marked reduction in Yki phosphorylation without affecting the Yki/Wts interaction. We suggest that Kibra is part of an apical scaffold that promotes SWH pathway activity.
Copyright (c) 2010 Elsevier Inc. All rights reserved.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Animals, Genetically Modified
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Apoptosis
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Base Sequence
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Cell Cycle Proteins / genetics
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Cell Cycle Proteins / physiology*
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Cell Proliferation
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Drosophila / genetics
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Drosophila / growth & development*
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Drosophila / physiology*
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Drosophila Proteins / antagonists & inhibitors
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Drosophila Proteins / genetics
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Drosophila Proteins / physiology*
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Epistasis, Genetic
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Female
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Gene Expression Regulation, Developmental
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Humans
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Intracellular Signaling Peptides and Proteins / genetics
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Intracellular Signaling Peptides and Proteins / physiology*
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Membrane Proteins / genetics
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Membrane Proteins / physiology
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Neurofibromin 2 / genetics
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Neurofibromin 2 / physiology
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Ovarian Follicle / cytology
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Ovarian Follicle / physiology
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Phenotype
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Protein Kinases / genetics
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Protein Kinases / physiology*
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Protein Serine-Threonine Kinases / genetics
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Protein Serine-Threonine Kinases / physiology*
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RNA Interference
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Signal Transduction
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Tumor Suppressor Proteins / antagonists & inhibitors
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Tumor Suppressor Proteins / genetics
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Tumor Suppressor Proteins / physiology*
Substances
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Cell Cycle Proteins
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Drosophila Proteins
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Intracellular Signaling Peptides and Proteins
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Membrane Proteins
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Neurofibromin 2
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Tumor Suppressor Proteins
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ex protein, Drosophila
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kibra protein, Drosophila
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merlin, Drosophila
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sav protein, Drosophila
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Protein Kinases
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wts protein, Drosophila
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Protein Serine-Threonine Kinases
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hpo protein, Drosophila