Abstract
Overexpression of phosphatase of regenerating liver (PRL)-3 is associated with the progression of diverse human cancers. We show that the overexpression of PRL-3 protein is not directly associated with its transcript levels, indicating the existence of an underlying posttranscriptional regulation. The 5' untranslanted region (UTR) of PRL-3 mRNA possesses triple GCCCAG motifs capable of suppressing mRNA translation through interaction with PolyC-RNA-binding protein 1 (PCBP1), which retards PRL-3 mRNA transcript incorporation into polyribosomes. Overexpression of PCBP1 inhibits PRL-3 expression and inactivates AKT, whereas knockdown of PCBP1 causes upregulation of PRL-3 protein levels, activation of AKT, and promotion of tumorigenesis. An inverse correlation between protein levels of PRL-3 and PCBP1 in human primary cancers supports the clinical relevance.
Copyright (c) 2010 Elsevier Inc. All rights reserved.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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5' Untranslated Regions / genetics
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Animals
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Blotting, Western
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Cell Line, Tumor
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DNA-Binding Proteins
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Electrophoretic Mobility Shift Assay
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Gene Expression Regulation, Neoplastic
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Heterogeneous-Nuclear Ribonucleoproteins / genetics
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Heterogeneous-Nuclear Ribonucleoproteins / metabolism*
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Humans
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Immunoenzyme Techniques
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Luciferases / metabolism
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Lymphatic Metastasis
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Mice
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Mice, Nude
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Neoplasm Metastasis
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Neoplasm Proteins / antagonists & inhibitors
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Neoplasm Proteins / genetics*
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Neoplasm Proteins / metabolism
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Neoplasms / genetics
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Neoplasms / metabolism*
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Neoplasms / pathology
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Polyribosomes / metabolism
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Promoter Regions, Genetic
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Protein Biosynthesis*
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Protein Tyrosine Phosphatases / antagonists & inhibitors
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Protein Tyrosine Phosphatases / genetics*
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Protein Tyrosine Phosphatases / metabolism
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Proto-Oncogene Proteins c-akt / genetics
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Proto-Oncogene Proteins c-akt / metabolism
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RNA, Messenger / genetics
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RNA, Small Interfering / pharmacology
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RNA-Binding Proteins
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Reverse Transcriptase Polymerase Chain Reaction
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Spectrometry, Mass, Matrix-Assisted Laser Desorption-Ionization
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Tissue Array Analysis
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Xenograft Model Antitumor Assays
Substances
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5' Untranslated Regions
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DNA-Binding Proteins
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Heterogeneous-Nuclear Ribonucleoproteins
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Neoplasm Proteins
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PCBP1 protein, human
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RNA, Messenger
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RNA, Small Interfering
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RNA-Binding Proteins
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Luciferases
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Proto-Oncogene Proteins c-akt
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PTP4A3 protein, human
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Protein Tyrosine Phosphatases