Activation of ephrin A proteins influences hematopoietic stem cell adhesion and trafficking patterns

Exp Hematol. 2010 Nov;38(11):1087-98. doi: 10.1016/j.exphem.2010.07.007. Epub 2010 Jul 23.

Abstract

Objective: To determine if Eph receptors and ephrins can modulate the homing of hematopoietic cells in a murine bone marrow transplantation model.

Materials and methods: EphA and ephrin A gene expression by mouse hematopoietic stem cells and the progenitor cell line FDCP-1 was determined by real-time reverse transcription polymerase chain reaction and flow cytometry. The effect of ephrin A activation on adhesion of hematopoietic progenitors was determined by in vitro adhesion assays in which cells were exposed to fibronectin or vascular cell adhesion molecule-1 (VCAM-1) and an increasing gradient of immobilized EphA3-Fc. Adhesion to fibronectin and VCAM-1 was further investigated using soluble preclustered EphA3-Fc. We used soluble unclustered EphA3-Fc as an antagonist to block endogenous EphA-ephrin A interactions in vivo. The effect of injecting soluble EphA3-Fc on the mobilization of hematopoietic progenitor cells was examined. We determined the effect on short-term homing by pretreating bone marrow cells with EphA3-Fc or the control IgG before infusion into lethally irradiated mice.

Results: Preclustered and immobilized EphA3-Fc increased adhesion of progenitor cells and FDCP-1 to fibronectin and VCAM-1 (1.6- to 2-fold higher adhesion; p < 0.05) relative to control (0 μ/cm(2) EphA3-Fc extracellular molecule alone). Injection of the antagonist soluble EphA3-Fc increased progenitor cell and colony-forming unit-spleen cells in the peripheral blood (42% greater colony-forming unit in culture; p < 0.05, 3.8-fold higher colony-forming unit-spleen) relative to control.

Conclusion: Treating bone marrow cells with EphA3-Fc resulted in a reduction by 31% in donor stem cells homing to the bone marrow and accumulation of donor cells in recipient spleens (50% greater than control) and greater recovery of donor stem cells from the peripheral blood.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Antibodies / immunology
  • Antibodies / pharmacology
  • Blotting, Western
  • Bone Marrow Cells / cytology
  • Bone Marrow Cells / drug effects
  • Bone Marrow Cells / metabolism*
  • Bone Marrow Transplantation
  • Cell Adhesion / drug effects
  • Cell Line
  • Cells, Cultured
  • Colony-Forming Units Assay
  • Ephrin-A3 / genetics
  • Ephrin-A3 / metabolism
  • Ephrins / genetics
  • Ephrins / metabolism*
  • Female
  • Fibronectins / metabolism
  • Flow Cytometry
  • Gene Expression
  • Hematopoietic Stem Cell Mobilization
  • Hematopoietic Stem Cells / cytology
  • Hematopoietic Stem Cells / metabolism*
  • Humans
  • Mice
  • Mice, Congenic
  • Mice, Inbred C57BL
  • Receptor, EphA3 / genetics
  • Receptor, EphA3 / immunology
  • Receptor, EphA3 / metabolism
  • Receptors, Eph Family / genetics
  • Receptors, Eph Family / immunology
  • Receptors, Eph Family / metabolism*
  • Reverse Transcriptase Polymerase Chain Reaction
  • Stromal Cells / cytology
  • Stromal Cells / metabolism
  • Vascular Cell Adhesion Molecule-1 / metabolism

Substances

  • Antibodies
  • Ephrin-A3
  • Ephrins
  • Fibronectins
  • Vascular Cell Adhesion Molecule-1
  • Receptor, EphA3
  • Receptors, Eph Family