Variation in FGF1, FOXE1, and TIMP2 genes is associated with nonsyndromic cleft lip with or without cleft palate

Birth Defects Res A Clin Mol Teratol. 2011 Apr;91(4):218-25. doi: 10.1002/bdra.20791. Epub 2011 Apr 1.

Abstract

Background: Nonsyndromic cleft lip with or without cleft palate (CL/P) is a common complex birth defect caused by the interaction between multiple genes and environmental factors.

Methods: Five hundred and eighty-seven single nucleotide polymorphisms in 40 candidate genes related to orofacial clefting were tested for association with CL/P in a clefting sample composed of 300 patients and 606 controls from Estonian, Latvian, and Lithuanian populations.

Results: In case-control comparisons, the minor alleles of FGF1 rs34010 (p = 4.56 × 10(-4) ), WNT9B rs4968282 (p = 0.0013), and FOXE1 rs7860144 (p = 0.0021) were associated with a decreased risk of CL/P. Multiple haplotypes in FGF1, FOXE1, and TIMP2 and haplotypes in WNT9B, PVRL2, and LHX8 were associated with CL/P. The strongest association was found for protective haplotype rs250092/rs34010 GT in the FGF1 gene (p = 5.01 × 10(-4) ). The strongest epistatic interaction was observed between the COL2A1 and WNT3 genes.

Conclusions: Our results provide for the first time evidence implicating FGF1 in the occurrence of CL/P, and support TIMP2 and WNT9B as novel loci predisposing to CL/P. We have also replicated recently reported significant associations between variants in or near FOXE1 and CL/P. It is likely that variation in FOXE1, TIMP2, and the FGF and Wnt signaling pathway genes confers susceptibility to nonsyndromic CL/P in Northeastern European populations.

MeSH terms

  • Case-Control Studies
  • Cell Adhesion Molecules / genetics
  • Cleft Lip / epidemiology
  • Cleft Lip / genetics*
  • Cleft Palate / epidemiology
  • Cleft Palate / genetics*
  • Epistasis, Genetic
  • Estonia / epidemiology
  • Female
  • Fibroblast Growth Factor 1 / genetics*
  • Forkhead Transcription Factors / genetics*
  • Genetic Loci
  • Haplotypes
  • Homeodomain Proteins / genetics
  • Humans
  • LIM-Homeodomain Proteins
  • Latvia / epidemiology
  • Lithuania / epidemiology
  • Male
  • Nectins
  • Polymorphism, Single Nucleotide*
  • Signal Transduction
  • Tissue Inhibitor of Metalloproteinase-2 / genetics*
  • Transcription Factors
  • Wnt Proteins / genetics

Substances

  • Cell Adhesion Molecules
  • FOXE1 protein, human
  • Forkhead Transcription Factors
  • Homeodomain Proteins
  • LIM homeobox protein 8
  • LIM-Homeodomain Proteins
  • Nectins
  • Transcription Factors
  • WNT9B protein, human
  • Wnt Proteins
  • Fibroblast Growth Factor 1
  • Tissue Inhibitor of Metalloproteinase-2