Pseudo-wild type revertants from inactive apocytochrome b mutants as a tool for the analysis of the structure/function relationships of the mitochondrial ubiquinol-cytochrome c reductase of Saccharomyces cerevisiae

J Biol Chem. 1990 Feb 25;265(6):3332-9.

Abstract

We have analyzed the structure/function relationships of the yeast mitochondrial cytochrome b with a new methodology based upon the isolation of pseudo-wild type revertants from well-characterized cytochrome b respiratory deficient mutants. Our goal was to determine how cytochrome b function could be restored in such mutants, at least to some degree, by suppressor mutations within the protein. True wild type revertants were differentiated from pseudo-wild type revertants by the use of a simple and rapid screening technique based upon oligonucleotide hybridization. This can easily be used to analyze a large number of revertants. The suppressor mutations responsible for the restoration of respiratory competence were identified by sequencing the revertant's cytochrome b mRNA in crude mitochondrial RNA preparations. Using this new method we have analyzed 210 independent revertants. We report here nine novel cytochrome b structures conferring a variety of respiratory sufficient phenotypes, obtained from five respiratory deficient mutations affecting a short region of the protein (positions 131-138 of the polypeptide chain), presumably belonging to the ubiquinol oxidizing center of the bc1 complex.

Publication types

  • Comparative Study
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Amino Acid Sequence
  • Apoproteins / genetics*
  • Apoproteins / metabolism
  • Base Sequence
  • Cytochrome b Group / genetics*
  • Cytochrome b Group / metabolism
  • Cytochromes b
  • Electron Transport Complex III / metabolism*
  • Genes, Fungal
  • Intracellular Membranes / metabolism
  • Mitochondria / metabolism*
  • Molecular Sequence Data
  • Mutation*
  • Nucleic Acid Hybridization
  • Oligonucleotide Probes
  • Protein Conformation
  • RNA, Fungal / genetics
  • Saccharomyces cerevisiae / genetics*
  • Saccharomyces cerevisiae / growth & development
  • Saccharomyces cerevisiae / metabolism
  • Sequence Homology, Nucleic Acid
  • Submitochondrial Particles / metabolism

Substances

  • Apoproteins
  • Cytochrome b Group
  • Oligonucleotide Probes
  • RNA, Fungal
  • Cytochromes b
  • Electron Transport Complex III