Abstract
Hemophagocytic lymphohistiocytosis (HLH) manifests as the uncontrolled activation of T lymphocytes and macrophages infiltrating multiple organs. Molecular studies of individuals with HLH have demonstrated in most of these conditions a critical role of granule-dependent cytotoxic activity in the regulation of lymphocyte homeostasis, and have allowed the characterization of key effectors regulating cytotoxic granule release. The cytolytic process may now be considered a multistep process, including cell activation; the polarization of cytotoxic granules toward the conjugated target cell; the tethering, priming, and fusion of the cytotoxic granules with the plasma membrane; and the release of their contents (perforin and granzymes) into the intercellular cleft, leading to target cell death. Cytolytic cells have a second effector function involving the production of cytokines, principally γ-interferon, which is secreted independently of the exocytosis cytotoxic granule pathway. An analysis of the mechanisms underlying HLH has identified γ-interferon as a key cytokine inducing uncontrolled macrophage activation, and thus represents a potential therapeutic target.
© 2011 New York Academy of Sciences.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Cytoplasmic Granules / immunology
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Cytoplasmic Granules / metabolism
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Disease Models, Animal
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Humans
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Interferon-gamma / genetics
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Interferon-gamma / metabolism
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Intracellular Signaling Peptides and Proteins / genetics
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Intracellular Signaling Peptides and Proteins / metabolism
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LIM Domain Proteins / genetics
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LIM Domain Proteins / metabolism
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LIM-Homeodomain Proteins / genetics
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LIM-Homeodomain Proteins / metabolism
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Lymphohistiocytosis, Hemophagocytic / genetics*
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Lymphohistiocytosis, Hemophagocytic / immunology*
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Membrane Proteins / genetics
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Membrane Proteins / metabolism
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Mice
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Munc18 Proteins / genetics
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Munc18 Proteins / metabolism
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Muscle Proteins / genetics
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Muscle Proteins / metabolism
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Perforin / genetics
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Perforin / metabolism
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Qa-SNARE Proteins / genetics
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Qa-SNARE Proteins / metabolism
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T-Lymphocytes, Cytotoxic / immunology
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Transcription Factors / genetics
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Transcription Factors / metabolism
Substances
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FHL2 protein, human
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FHL3 protein, human
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FHL5 protein, human
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Intracellular Signaling Peptides and Proteins
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LIM Domain Proteins
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LIM-Homeodomain Proteins
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Membrane Proteins
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Munc18 Proteins
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Muscle Proteins
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Qa-SNARE Proteins
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STX11 protein, human
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STXBP2 protein, human
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Transcription Factors
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UNC13D protein, human
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Perforin
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Interferon-gamma