Distinct TLR-mediated pathways regulate house dust mite-induced allergic disease in the upper and lower airways

J Allergy Clin Immunol. 2013 Feb;131(2):549-61. doi: 10.1016/j.jaci.2012.07.050. Epub 2012 Oct 1.

Abstract

Background: Allergic rhinitis (AR) and asthma are 2 entities of allergic airway diseases that frequently occur together, which is referred to as united airways. In contrast to this general concept, we hypothesized that innate immunity of the upper and lower airways is respectively distinctive, because the immunologic conditions of the nasal and lung mucosa as well as the functions of the immune cells within their epithelia are different.

Objective: We wanted to identify distinctive mechanisms of innate immunity in the nose and lung mucosa, which are responsible for house dust mite (HDM)-induced AR and allergic asthma (AA), respectively.

Methods: We constructed a mouse model of AR or AA induced by sensitization and consequent provocation with HDM extracts.

Results: HDM-derived β-glucans, rather than LPS, were proven to be essential to activating innate immunity in the nasal mucosa and triggering AR, which depended on Toll-like receptor 2 (TLR2), but not on TLR4; however, the LPS/TLR4 signaling axis, rather than β-glucans/TLR2, was critical to HDM-induced AA. These differences were attributed to the specific role of β-glucans and LPS in inducing the surface expression of TLR2 and TLR4 and their translocation to lipid rafts in nasal and bronchial epithelial cells, respectively. We also showed that dual oxidase 2-generated reactive oxygen species mediate both β-glucan-induced TLR2 activation and LPS-induced TLR4 activation.

Conclusions: We describe a novel finding of distinctive innate immunity of the nose and lungs, respectively, which trigger AR and AA, by showing the critical role of HDM-induced TLR activation via dual oxidase 2-mediated reactive oxygen species.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Asthma / immunology
  • Asthma / metabolism
  • Dual Oxidases
  • Epithelial Cells / immunology
  • Epithelial Cells / metabolism
  • Hypersensitivity / immunology*
  • Hypersensitivity / metabolism
  • Immunity, Innate / immunology
  • Lipopolysaccharides / immunology
  • Lung / immunology*
  • Lung / metabolism
  • Mice
  • NADPH Oxidases / immunology
  • NADPH Oxidases / metabolism
  • Nasal Mucosa / immunology*
  • Nasal Mucosa / metabolism
  • Pyroglyphidae / immunology*
  • Reactive Oxygen Species / immunology
  • Reactive Oxygen Species / metabolism
  • Respiratory Mucosa / immunology
  • Respiratory Mucosa / metabolism
  • Respiratory System / immunology*
  • Respiratory System / metabolism
  • Rhinitis, Allergic
  • Rhinitis, Allergic, Perennial / immunology
  • Rhinitis, Allergic, Perennial / metabolism
  • Toll-Like Receptor 2 / immunology
  • Toll-Like Receptor 2 / metabolism*
  • Toll-Like Receptor 4 / immunology
  • Toll-Like Receptor 4 / metabolism*
  • beta-Glucans / immunology
  • beta-Glucans / metabolism

Substances

  • Lipopolysaccharides
  • Reactive Oxygen Species
  • Tlr2 protein, mouse
  • Tlr4 protein, mouse
  • Toll-Like Receptor 2
  • Toll-Like Receptor 4
  • beta-Glucans
  • Dual Oxidases
  • NADPH Oxidases
  • Duox1 protein, mouse