1 Whitehead Institute for Biomedical Research, 9 Cambridge Center, Cambridge, MA 02142, USA.
2 Whitehead Institute for Biomedical Research, 9 Cambridge Center, Cambridge, MA 02142, USA ; Harvard-MIT Division of Health Sciences and Technology, MIT, Cambridge, MA 02139, USA ; Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA 02142, USA ; Department of Molecular Biology and Diabetes Unit, Massachusetts General Hospital, Boston, MA 02114, USA.