The octadecaneuropeptide ODN prevents 6-hydroxydopamine-induced apoptosis of cerebellar granule neurons through a PKC-MAPK-dependent pathway

J Neurochem. 2013 May;125(4):620-33. doi: 10.1111/jnc.12140. Epub 2013 Feb 19.

Abstract

Oxidative stress, induced by various neurodegenerative diseases, initiates a cascade of events leading to apoptosis, and thus plays a critical role in neuronal injury. In this study, we have investigated the potential neuroprotective effect of the octadecaneuropeptide (ODN) on 6-hydroxydopamine (6-OHDA)-induced oxidative stress and apoptosis in cerebellar granule neurons (CGN). ODN, which is produced by astrocytes, is an endogenous ligand for both central-type benzodiazepine receptors (CBR) and a metabotropic receptor. Incubation of neurons with subnanomolar concentrations of ODN (10⁻¹⁸ to 10⁻¹² M) inhibited 6-OHDA-evoked cell death in a concentration-dependent manner. The effect of ODN on neuronal survival was abrogated by the metabotropic receptor antagonist, cyclo₁₋₈ [DLeu⁵]OP, but not by a CBR antagonist. ODN stimulated polyphosphoinositide turnover and ERK phosphorylation in CGN. The protective effect of ODN against 6-OHDA toxicity involved the phospholipase C/ERK MAPK transduction cascade. 6-OHDA treatment induced an accumulation of reactive oxygen species, an increase of the expression of the pro-apoptotic gene Bax, a drop of the mitochondrial membrane potential and a stimulation of caspase-3 activity. Exposure of 6-OHDA-treated cells to ODN blocked all the deleterious effects of the toxin. Taken together, these data demonstrate for the first time that ODN is a neuroprotective agent that prevents 6-OHDA-induced oxidative stress and apoptotic cell death.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Apoptosis / drug effects*
  • Apoptosis / physiology
  • Caspase 3 / metabolism
  • Cerebellum / drug effects*
  • Cerebellum / pathology
  • Cyclin D1 / genetics
  • Cyclin D1 / metabolism
  • Diazepam Binding Inhibitor / pharmacology*
  • Glutathione / metabolism
  • MAP Kinase Signaling System / drug effects*
  • MAP Kinase Signaling System / physiology
  • Neurons / drug effects*
  • Neurons / pathology
  • Neuropeptides / pharmacology*
  • Neuroprotective Agents / pharmacology
  • Oxidative Stress / physiology
  • Oxidopamine / toxicity*
  • Peptide Fragments / pharmacology*
  • Phosphorylation / drug effects
  • Phosphorylation / physiology
  • Protein Kinase C / metabolism*
  • Rats
  • Rats, Wistar
  • Reactive Oxygen Species / metabolism
  • Sympatholytics / toxicity

Substances

  • Diazepam Binding Inhibitor
  • Neuropeptides
  • Neuroprotective Agents
  • Peptide Fragments
  • Reactive Oxygen Species
  • Sympatholytics
  • diazepam binding inhibitor (33-50)
  • Cyclin D1
  • Oxidopamine
  • Protein Kinase C
  • Casp3 protein, rat
  • Caspase 3
  • Glutathione