Regulation of endoplasmic reticulum stress in rat cortex by p62/ZIP through the Keap1-Nrf2-ARE signalling pathway after transient focal cerebral ischaemia

Brain Inj. 2013;27(7-8):924-33. doi: 10.3109/02699052.2013.793397. Epub 2013 Jun 19.

Abstract

Primary objective: p62/ZIP as the autophagy receptor can transport the misfolded proteins to a macroautophagy-lysosome system for degradation and also create a positive feedback loop between p62/ZIP and Nrf2. However, the role of p62/ZIP on cerebral ischaemia is unclear. The aim of this study was to evaluate the role of p62/ZIP in the regulation of endoplasmic reticulum(ER) stress induced by cerebral ischaemia/reperfusion.

Research design: Different ischemic periods were designed by transient middle cerebral artery occlusion (tMCAO) using the suture method.

Methods and procedures: At 24 hours after reperfusion, the ischaemic brain tissue was studied histologically and biochemically for autophagic, ER stress and Keap1-Nrf2-ARE signalling pathway markers.

Main outcomes and results: Prolongation of ischaemia significantly increased the cortical injury observed in rats and was associated with a gradual increase in the protein expression of ubiquitin-aggregates, Grp78, GADD153/CHOP and p62/ZIP. Autophagy marker Atg12-Atg5 and LC3-PE increased and then decreased. Moreover, p62/ZIP mRNA expression increased and then decreased and was consistent with Nrf2 activation.

Conclusions: p62/ZIP not only plays a key role in scavenging protein aggregates during autophagy, but it may also be involved in preventing oxidative injury and alleviating ER stress through the Keap1-Nrf2-ARE signalling pathway during cerebral ischaemia/reperfusion injury.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Autophagy
  • Blotting, Western
  • Brain / blood supply
  • Brain / metabolism
  • Brain Ischemia* / metabolism
  • Endoplasmic Reticulum Stress*
  • Heat-Shock Proteins* / metabolism
  • Immunohistochemistry
  • Intracellular Signaling Peptides and Proteins* / metabolism
  • Kelch-Like ECH-Associated Protein 1
  • Male
  • NF-E2-Related Factor 2 / metabolism
  • Rats
  • Rats, Wistar
  • Reperfusion Injury* / metabolism
  • Reperfusion Injury* / pathology
  • Sequestosome-1 Protein
  • Signal Transduction*
  • Time Factors
  • Transcription Factor CHOP / metabolism

Substances

  • Ddit3 protein, rat
  • GRP78 protein, rat
  • Heat-Shock Proteins
  • Intracellular Signaling Peptides and Proteins
  • KEAP1 protein, rat
  • Kelch-Like ECH-Associated Protein 1
  • Sequestosome-1 Protein
  • Sqstm1 protein, rat
  • Transcription Factor CHOP
  • NF-E2-Related Factor 2