The role of Y84 on domain 1 and Y87 on domain 2 of Paragonimus westermani taurocyamine kinase: Insights on the substrate binding mechanism of a trematode phosphagen kinase

Exp Parasitol. 2013 Dec;135(4):695-700. doi: 10.1016/j.exppara.2013.10.008. Epub 2013 Oct 30.

Abstract

The two-domain taurocyamine kinase (TK) from Paragonimus westermani was suggested to have a unique substrate binding mechanism. We performed site-directed mutagenesis on each domain of this TK and compared the kinetic parameters Km(Tc) and Vmax with that of the wild-type to determine putative amino acids involved in substrate recognition and binding. Replacement of Y84 on domain 1 and Y87 on domain 2 with R resulted in the loss of activity for the substrate taurocyamine. Y84E mutant has a dramatic decrease in affinity and activity for taurocyamine while Y87E has completely lost catalytic activity. Substituting H and I on the said positions also resulted in significant changes in activity. Mutation of the residues A59 on the GS region of domain 1 also caused significant decrease in affinity and activity while mutation on the equivalent position on domain 2 resulted in complete loss of activity.

Keywords: AK; CK; GK; LK; PK; Paragonimus westermani; Phosphagen kinase; Site-directed mutagenesis; TK; Taurocyamine kinase; Trematode; arginine kinase; creatine kinase; glycocyamine kinase; lombricine kinase; phosphagen kinase; taurocyamine kinase.

MeSH terms

  • Amino Acid Sequence
  • Animals
  • Base Sequence
  • Conserved Sequence
  • Molecular Sequence Data
  • Mutagenesis, Site-Directed
  • Paragonimus westermani / enzymology*
  • Phosphotransferases (Nitrogenous Group Acceptor) / chemistry
  • Phosphotransferases (Nitrogenous Group Acceptor) / genetics
  • Phosphotransferases (Nitrogenous Group Acceptor) / metabolism*
  • Protein Structure, Tertiary* / genetics
  • Sequence Alignment
  • Substrate Specificity
  • Taurine / analogs & derivatives*
  • Taurine / metabolism
  • Tyrosine* / chemistry
  • Tyrosine* / genetics

Substances

  • Taurine
  • Tyrosine
  • taurocyamine
  • Phosphotransferases (Nitrogenous Group Acceptor)
  • taurocyamine kinase