Downregulation of microRNA-24 and -181 parallels the upregulation of IFN-γ secreted by activated human CD4 lymphocytes

Hum Immunol. 2014 Jul;75(7):677-85. doi: 10.1016/j.humimm.2014.01.007. Epub 2014 Apr 3.

Abstract

IFN-γ is a cytokine with important roles in the innate and adaptive immune responses. This cytokine is secreted by activated T cells, NK cells and macrophages. Studies on the regulation of human IFN-γ expression had been previously focused on the promoter region. Consequently, the role of microRNAs (miRs) in this regulation has not been investigated yet. As miR-24 and miR-181 were found to have potential target sites in IFN-γ mRNA 3'UTR, we assessed their impact on IFN-γ expression by co-stimulating PB CD4+ T cells with anti-CD3, anti-CD28, IL-12, and IL-18. This co-stimulation cocktail induced an abundant secretion of IFN-γ together with a down-regulation of miR-24, and miR-181. Existence of a link between these two phenomena was further substantiated by transfection and transduction assays that showed that these two miRs negatively regulate IFN-γ expression by directly binding to their target sites in the mRNA. Thus, identifying target sites for miR-24 and miR-181 in IFN-γ-3'UTR points to a novel regulatory mechanism of this crucial gene.

Keywords: Activated CD4 T lymphocyte; IFN-γ; microRNA.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • 3' Untranslated Regions*
  • Antibodies / pharmacology
  • Base Sequence
  • Binding Sites
  • CD28 Antigens / antagonists & inhibitors
  • CD28 Antigens / genetics
  • CD28 Antigens / immunology
  • CD3 Complex / genetics
  • CD3 Complex / immunology
  • CD4-Positive T-Lymphocytes / cytology
  • CD4-Positive T-Lymphocytes / drug effects
  • CD4-Positive T-Lymphocytes / immunology*
  • Cell Separation
  • Gene Expression Regulation*
  • Humans
  • Interferon-gamma / biosynthesis
  • Interferon-gamma / genetics*
  • Interferon-gamma / metabolism
  • Interleukin-12 / pharmacology
  • Interleukin-18 / pharmacology
  • Lymphocyte Activation / drug effects
  • MicroRNAs / antagonists & inhibitors
  • MicroRNAs / genetics*
  • MicroRNAs / immunology
  • Molecular Sequence Data
  • Primary Cell Culture
  • Signal Transduction

Substances

  • 3' Untranslated Regions
  • Antibodies
  • CD28 Antigens
  • CD3 Complex
  • IL18 protein, human
  • Interleukin-18
  • MIRN24 microRNA, human
  • MIrn181 microRNA, human
  • MicroRNAs
  • Interleukin-12
  • Interferon-gamma