Lyn sustains oncogenic signaling in chronic lymphocytic leukemia by strengthening SET-mediated inhibition of PP2A

Blood. 2015 Jun 11;125(24):3747-55. doi: 10.1182/blood-2014-12-619155. Epub 2015 Apr 30.

Abstract

Aberrant protein kinase activities, and the consequent dramatic increase of Ser/Thr and -Tyr phosphorylation, promote the deregulation of the survival pathways in chronic lymphocytic leukemia (CLL), which is crucial to the pathogenesis and progression of the disease. In this study, we show that the tumor suppressor protein phosphatase 2A (PP2A), one of the major Ser/Thr phosphatases, is in an inhibited form because of the synergistic contribution of 2 events, the interaction with its physiologic inhibitor SET and the phosphorylation of Y307 of the catalytic subunit of PP2A. The latter event is mediated by Lyn, a Src family kinase previously found to be overexpressed, delocalized, and constitutively active in CLL cells. This Lyn/PP2A axis accounts for the persistent high level of phosphorylation of the phosphatase's targets and represents a key connection linking phosphotyrosine- and phosphoserine/threonine-mediated oncogenic signals. The data herein presented show that the disruption of the SET/PP2A complex by a novel FTY720-analog (MP07-66) devoid of immunosuppressive effects leads to the reactivation of PP2A, which in turn triggers apoptosis of CLL cells. When used in combination with SFK inhibitors, the action of MP07-66 is synergistically amplified, providing a new option in the therapeutic strategy for CLL patients.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Apoptosis / drug effects
  • DNA-Binding Proteins
  • Fingolimod Hydrochloride
  • HSP90 Heat-Shock Proteins / metabolism
  • Histone Chaperones / metabolism*
  • Humans
  • Immunosuppressive Agents / chemistry
  • Immunosuppressive Agents / pharmacology*
  • Leukemia, Lymphocytic, Chronic, B-Cell / drug therapy*
  • Leukemia, Lymphocytic, Chronic, B-Cell / metabolism*
  • Phosphorylation / drug effects
  • Phosphotyrosine / metabolism
  • Propylene Glycols / chemistry
  • Propylene Glycols / pharmacology*
  • Protein Interaction Maps / drug effects
  • Protein Phosphatase 2 / metabolism*
  • Signal Transduction / drug effects
  • Sphingosine / analogs & derivatives*
  • Sphingosine / chemistry
  • Sphingosine / pharmacology
  • Transcription Factors / metabolism*
  • Tumor Cells, Cultured
  • src-Family Kinases / metabolism*

Substances

  • DNA-Binding Proteins
  • HSP90 Heat-Shock Proteins
  • Histone Chaperones
  • Immunosuppressive Agents
  • Propylene Glycols
  • SET protein, human
  • Transcription Factors
  • Phosphotyrosine
  • lyn protein-tyrosine kinase
  • src-Family Kinases
  • Protein Phosphatase 2
  • Fingolimod Hydrochloride
  • Sphingosine