Lack of mature lymphocytes results in obese but metabolically healthy mice when fed a high-fat diet

Int J Obes (Lond). 2015 Oct;39(10):1548-57. doi: 10.1038/ijo.2015.93. Epub 2015 May 21.

Abstract

Background/objectives: Obesity is characterized by chronic inflammation and immune dysregulation, as well as insulin resistance, but the link between obesity and adaptive immunity remains to be fully studied.

Methods: To elucidate the role of adaptive immunity on body composition, glucose homeostasis and inflammation, recombination-activating gene 1 knockout (Rag1-/-) mice, without mature T-lymphocytes or B-lymphocytes, were maintained on a low- or high-fat diet (LFD and HFD, respectively) for 11 weeks.

Results: Rag1-/- mice fed HFD gained significantly more weight and had increased body fat compared with wild type. Downregulation of energy expenditure as well as brown fat uncoupling protein UCP-1 and UCP-3 gene expression were noticed in HFD-fed Rag1-/- mice compared with LFD. HFD mice had significantly decreased energy intake compared with LFD mice, consistent with decreased agouti-related protein and increased pro-opiomelanocortin gene expression levels in the hypothalamus. Moreover, compared with wild type, Rag1-/- mice had lower interleukin (IL)-4 levels, a cytokine recently found to induce browning in white adipocytes, and higher IL-12 levels in HFD-fed Rag1-/- mice. Despite that HFD Rag1-/- mice were more obese, they had similar glucose, insulin and adiponectin levels, while leptin was marginally increased.

Conclusions: Mice with deficiency in adaptive immunity are obese, partly owing to decreased energy expenditure, but are metabolically normal, suggesting that mature lymphocytes have necessary roles in the development of obesity-related metabolic dysregulation.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't
  • Research Support, U.S. Gov't, Non-P.H.S.

MeSH terms

  • Adaptive Immunity
  • Animals
  • Diet, High-Fat
  • Disease Models, Animal
  • Homeodomain Proteins / metabolism*
  • Hypothalamus / pathology*
  • Inflammation
  • Insulin / metabolism*
  • Insulin Resistance
  • Lymphocytes / metabolism*
  • Male
  • Mice
  • Mice, Inbred C57BL
  • Obesity / metabolism
  • Obesity / pathology*

Substances

  • Homeodomain Proteins
  • Insulin
  • RAG-1 protein