Abstract
Detection of intracellular DNA triggers activation of the stimulator of IFN genes-dependent IFN-stimulatory DNA (ISD) pathway, which is essential for antiviral immune responses. However, chronic activation of this pathway is implicated in autoimmunity. Mutations in TREX1, a 3' repair exonuclease that degrades cytosolic DNA, cause Aicardi-Goutières syndrome and chilblain lupus. Trex1 (-/-) mice develop lethal, IFN-driven autoimmune disease that is dependent on activation of the ISD pathway, but the DNA sensors that detect the endogenous DNA that accumulates in Trex1 (-/-) mice have not been defined. Multiple DNA sensors have been proposed to activate the ISD pathway, including cyclic GMP-AMP synthase (cGAS). In this study, we show that Trex1 (-/-) mice lacking cGAS are completely protected from lethality, exhibit dramatically reduced tissue inflammation, and fail to develop autoantibodies. These findings implicate cGAS as a key driver of autoimmune disease and suggest that cGAS inhibitors may be useful therapeutics for Aicardi-Goutières syndrome and related autoimmune diseases.
Copyright © 2015 by The American Association of Immunologists, Inc.
Publication types
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Research Support, N.I.H., Extramural
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Autoantibodies / blood
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Autoantibodies / immunology
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Autoimmune Diseases of the Nervous System / genetics
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Autoimmune Diseases of the Nervous System / immunology*
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Autoimmune Diseases of the Nervous System / metabolism
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Cells, Cultured
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Disease Models, Animal
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Embryo, Mammalian / cytology
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Exodeoxyribonucleases / genetics
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Exodeoxyribonucleases / immunology*
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Exodeoxyribonucleases / metabolism
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Fibroblasts / drug effects
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Fibroblasts / immunology
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Fibroblasts / metabolism
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Gene Expression / drug effects
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Gene Expression / immunology
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Humans
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Immunoblotting
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Inflammation / genetics
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Inflammation / immunology
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Inflammation / metabolism
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Interferon-beta / genetics
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Interferon-beta / immunology
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Interferon-beta / metabolism
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Interferons / immunology
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Interferons / pharmacology
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Macrophages / drug effects
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Macrophages / immunology
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Macrophages / metabolism
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Mice, Inbred C57BL
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Mice, Knockout
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Nervous System Malformations / genetics
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Nervous System Malformations / immunology*
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Nervous System Malformations / metabolism
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Nucleotides, Cyclic / immunology
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Nucleotides, Cyclic / metabolism
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Nucleotidyltransferases / genetics
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Nucleotidyltransferases / immunology*
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Nucleotidyltransferases / metabolism
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Phosphoproteins / genetics
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Phosphoproteins / immunology*
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Phosphoproteins / metabolism
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Reverse Transcriptase Polymerase Chain Reaction
Substances
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Autoantibodies
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Nucleotides, Cyclic
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Phosphoproteins
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cyclic guanosine monophosphate-adenosine monophosphate
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Interferon-beta
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Interferons
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Nucleotidyltransferases
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cGAS protein, mouse
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Exodeoxyribonucleases
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three prime repair exonuclease 1
Supplementary concepts
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Aicardi-Goutieres syndrome