Interaction of Heat Shock Protein Cpn10 with the Cyclin E/Cdk2 Substrate Nuclear Protein Ataxia-Telangiectasia (NPAT) Is Involved in Regulating Histone Transcription

J Biol Chem. 2015 Dec 4;290(49):29290-300. doi: 10.1074/jbc.M115.659201. Epub 2015 Oct 1.

Abstract

Precise modulation of histone gene transcription is critical for cell cycle progression. As a direct substrate of Cyclin E/CDK2, nuclear protein ataxia-telangiectasia (NPAT) is a crucial factor in regulating histone transcription and cell cycle progression. Here we identified that Cpn10/HSPE, a 10-kDa heat shock protein, is a novel interacting partner of NPAT. A pool of Cpn10 is colocalized with NPAT foci during G1 and S phases in nuclei. Gain- and loss-of-function experiments unraveled an essential role of Cpn10 in histone transcription. A conserved DLFD motif within Cpn10 was critical for targeting NPAT and modulating histone transcription. More importantly, knockdown of Cpn10 disrupted the focus formation of both NPAT and FADD-like interleukin-1β-converting enzyme-associated huge protein without affecting Coilin-positive Cajal bodies. Finally, Cpn10 is important for S phase progression and cell proliferation. Taken together, our finding revealed a novel role of Cpn10 in the spatial regulation of NPAT signaling and disclosed a previously unappreciated link between the heat shock protein and histone transcription regulation.

Keywords: Cpn10; NPAT; cell cycle; cell proliferation; cyclin; heat shock protein (HSP); histone; histone transcription.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Amino Acid Motifs
  • Cell Cycle
  • Cell Cycle Proteins / metabolism*
  • Cell Nucleus / metabolism
  • Cell Proliferation
  • Chaperonin 10 / metabolism*
  • Cyclin E / metabolism*
  • Cyclin-Dependent Kinase 2 / metabolism*
  • Disease Progression
  • HeLa Cells
  • Histones / metabolism*
  • Humans
  • Microscopy, Fluorescence
  • Nuclear Proteins / metabolism*
  • Pregnancy Proteins / metabolism*
  • RNA Interference
  • Signal Transduction
  • Suppressor Factors, Immunologic / metabolism*
  • Transcription, Genetic
  • Two-Hybrid System Techniques

Substances

  • Cell Cycle Proteins
  • Chaperonin 10
  • Cyclin E
  • Histones
  • NPAT protein, human
  • Nuclear Proteins
  • Pregnancy Proteins
  • Suppressor Factors, Immunologic
  • early pregnancy factor
  • CDK2 protein, human
  • Cyclin-Dependent Kinase 2