Type I IFN promotes NK cell expansion during viral infection by protecting NK cells against fratricide

J Exp Med. 2016 Feb 8;213(2):225-33. doi: 10.1084/jem.20150712. Epub 2016 Jan 11.

Abstract

Type I interferon (IFN) is crucial in host antiviral defense. Previous studies have described the pleiotropic role of type I IFNs on innate and adaptive immune cells during viral infection. Here, we demonstrate that natural killer (NK) cells from mice lacking the type I IFN-α receptor (Ifnar(-/-)) or STAT1 (which signals downstream of IFNAR) are defective in expansion and memory cell formation after mouse cytomegalovirus (MCMV) infection. Despite comparable proliferation, Ifnar(-/-) NK cells showed diminished protection against MCMV infection and exhibited more apoptosis compared with wild-type NK cells. Furthermore, we show that Ifnar(-/-) NK cells express increased levels of NK group 2 member D (NKG2D) ligands during viral infection and are susceptible to NK cell-mediated fratricide in a perforin- and NKG2D-dependent manner. Adoptive transfer of Ifnar(-/-) NK cells into NK cell-deficient mice reverses the defect in survival and expansion. Our study reveals a novel type I IFN-dependent mechanism by which NK cells evade mechanisms of cell death after viral infection.

Publication types

  • Research Support, N.I.H., Extramural

MeSH terms

  • Animals
  • Apoptosis
  • Cell Proliferation
  • Herpesviridae Infections / immunology*
  • Herpesviridae Infections / metabolism
  • Herpesviridae Infections / pathology
  • Interferon Type I / metabolism*
  • Killer Cells, Natural / immunology*
  • Killer Cells, Natural / metabolism
  • Killer Cells, Natural / pathology
  • Lymphocyte Activation
  • Mice
  • Mice, Inbred C57BL
  • Mice, Knockout
  • Muromegalovirus*
  • NK Cell Lectin-Like Receptor Subfamily K / metabolism
  • Perforin / deficiency
  • Perforin / genetics
  • Perforin / metabolism
  • Receptor, Interferon alpha-beta / deficiency
  • Receptor, Interferon alpha-beta / genetics
  • Receptor, Interferon alpha-beta / metabolism
  • STAT1 Transcription Factor / deficiency
  • STAT1 Transcription Factor / genetics
  • STAT1 Transcription Factor / metabolism

Substances

  • Ifnar1 protein, mouse
  • Interferon Type I
  • Klrk1 protein, mouse
  • NK Cell Lectin-Like Receptor Subfamily K
  • STAT1 Transcription Factor
  • Stat1 protein, mouse
  • perforin 1, mouse
  • Perforin
  • Receptor, Interferon alpha-beta