High-throughput FACS-based mutant screen identifies a gain-of-function allele of the Fusarium graminearum adenylyl cyclase causing deoxynivalenol over-production

Fungal Genet Biol. 2016 May:90:1-11. doi: 10.1016/j.fgb.2016.02.005. Epub 2016 Feb 27.

Abstract

Fusarium head blight and crown rot, caused by the fungal plant pathogen Fusarium graminearum, impose a major threat to global wheat production. During the infection, plants are contaminated with mycotoxins such as deoxynivalenol (DON), which can be toxic for humans and animals. In addition, DON is a major virulence factor during wheat infection. However, it is not fully understood how DON production is regulated in F. graminearum. In order to identify regulators of DON production, a high-throughput mutant screen using Fluorescence Activated Cell Sorting (FACS) of a mutagenised TRI5-GFP reporter strain was established and a mutant over-producing DON under repressive conditions identified. A gain-of-function mutation in the F. graminearum adenylyl cyclase (FAC1), which is a known positive regulator of DON production, was identified as the cause of this phenotype through genome sequencing and segregation analysis. Our results show that the high-throughput mutant screening procedure developed here can be applied for identification of fungal proteins involved in diverse processes.

Keywords: DON; Flow cytometry; Fluorescence activated cell sorting; Gibberella zeae; Trichothecene; cAMP.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adenosine Monophosphate / metabolism
  • Adenylyl Cyclases / genetics*
  • Adenylyl Cyclases / metabolism*
  • Alleles*
  • DNA, Fungal
  • Flow Cytometry / methods
  • Fungal Proteins / genetics
  • Fungal Proteins / metabolism
  • Fusarium / enzymology*
  • Fusarium / genetics*
  • Fusarium / metabolism
  • High-Throughput Screening Assays / methods
  • Mutation*
  • Mycotoxins / metabolism
  • Phenotype
  • Plant Diseases / microbiology
  • Trichothecenes / biosynthesis*
  • Trichothecenes / metabolism
  • Triticum / microbiology

Substances

  • DNA, Fungal
  • Fungal Proteins
  • Mycotoxins
  • Trichothecenes
  • Adenosine Monophosphate
  • Adenylyl Cyclases
  • deoxynivalenol