A potential contribution of antimicrobial peptide LL-37 to tissue fibrosis and vasculopathy in systemic sclerosis

Br J Dermatol. 2016 Dec;175(6):1195-1203. doi: 10.1111/bjd.14699. Epub 2016 Aug 30.

Abstract

Background: LL-37 is an antimicrobial peptide with pleiotropic effects on the immune system, angiogenesis and tissue remodelling. These are cardinal pathological events in systemic sclerosis (SSc).

Objectives: To elucidate the potential role of LL-37 in SSc.

Methods: The expression of target molecules was evaluated by immunostaining and quantitative reverse-transcription real-time polymerase chain reaction in human and murine skin. The mechanisms regulating LL-37 expression in endothelial cells were examined by gene silencing and chromatin immunoprecipitation. Serum LL-37 levels were determined by enzyme-linked immunosorbent assay.

Results: In SSc lesional skin, LL-37 expression was increased in dermal fibroblasts, perivascular inflammatory cells, keratinocytes and, particularly, dermal small vessels. Expression positively correlated with interferon-α expression, possibly reflecting LL-37-dependent induction of interferon-α. In SSc animal models, bleomycin-treated skin exhibited the expression pattern of CRAMP, a murine homologue of LL-37, similar to that of LL-37 in SSc lesional skin. Furthermore, Fli1+/- mice showed upregulated expression of CRAMP in dermal small vessels. Fli1 binding to the CAMP (LL-37 gene) promoter and Fli1 deficiency-dependent induction of LL-37 were also confirmed in human dermal microvascular endothelial cells. In the analysis of sera, patients with SSc had serum LL-37 levels significantly higher than in healthy controls. Furthermore, serum LL-37 levels positively correlated with skin score and the activity of alveolitis and were significantly elevated in patients with digital ulcers compared with those without.

Conclusions: LL-37 upregulation, induced by Fli1 deficiency at least in endothelial cells, potentially contributes to the development of skin sclerosis, interstitial lung disease and digital ulcers in SSc.

MeSH terms

  • Adult
  • Aged
  • Animals
  • Antimicrobial Cationic Peptides / metabolism
  • Cathelicidins / metabolism
  • Cathelicidins / physiology*
  • Endothelial Cells / metabolism
  • Female
  • Fibrosis / blood
  • Fibrosis / etiology
  • Humans
  • Interferon-alpha / metabolism
  • Mice, Inbred C57BL
  • Middle Aged
  • Proto-Oncogene Protein c-fli-1 / deficiency
  • Scleroderma, Systemic / blood
  • Scleroderma, Systemic / etiology*
  • Skin / pathology*
  • Up-Regulation / physiology
  • Vascular Diseases / blood
  • Vascular Diseases / etiology*

Substances

  • Antimicrobial Cationic Peptides
  • Cathelicidins
  • Fli1 protein, mouse
  • Interferon-alpha
  • Proto-Oncogene Protein c-fli-1