Rab27b is Involved in Lysosomal Exocytosis and Proteolipid Protein Trafficking in Oligodendrocytes

Neurosci Bull. 2016 Aug;32(4):331-40. doi: 10.1007/s12264-016-0045-6. Epub 2016 Jun 21.

Abstract

Myelination by oligodendrocytes in the central nervous system requires coordinated exocytosis and endocytosis of the major myelin protein, proteolipid protein (PLP). Here, we demonstrated that a small GTPase, Rab27b, is involved in PLP trafficking in oligodendrocytes. We showed that PLP co-localized with Rab27b in late endosomes/lysosomes in oligodendrocytes. Short hairpin-mediated knockdown of Rab27b not only reduced lysosomal exocytosis but also greatly diminished the surface expression of PLP in oligodendrocytes. In addition, knockdown of Rab27b reduced the myelin-like membranes induced by co-culture of oligodendrocytes and neurons. Our data suggest that Rab27b is involved in myelin biogenesis by regulating PLP transport from late endosomes/lysosomes to the cell membrane in oligodendrocytes.

Keywords: Lysosome; Myelination; Neuron-glial interaction; Oligodendrocyte; Rab27b.

MeSH terms

  • Animals
  • Animals, Newborn
  • Calcium Ionophores / pharmacology
  • Cathepsin D / metabolism
  • Cells, Cultured
  • Cerebral Cortex / cytology
  • Coculture Techniques
  • Embryo, Mammalian
  • Exocytosis / drug effects
  • Exocytosis / physiology*
  • Ionomycin / pharmacology
  • Lysosomes / drug effects
  • Lysosomes / metabolism*
  • Neurons / drug effects
  • Neurons / metabolism
  • Oligodendroglia / cytology*
  • Oligodendroglia / drug effects
  • Oligodendroglia / metabolism*
  • Protein Transport / drug effects
  • Proteolipids / metabolism*
  • RNA Interference / physiology
  • Rats
  • Rats, Sprague-Dawley
  • Time Factors
  • rab GTP-Binding Proteins / genetics
  • rab GTP-Binding Proteins / metabolism*

Substances

  • Calcium Ionophores
  • Proteolipids
  • Ionomycin
  • Cathepsin D
  • Rab27b protein, rat
  • rab GTP-Binding Proteins