Sequence conservation predicts T cell reactivity against ragweed allergens

Clin Exp Allergy. 2016 Sep;46(9):1194-205. doi: 10.1111/cea.12772. Epub 2016 Jul 26.

Abstract

Background: Ragweed is a major cause of seasonal allergy, affecting millions of people worldwide. Several allergens have been defined based on IgE reactivity, but their relative immunogenicity in terms of T cell responses has not been studied.

Objective: We comprehensively characterized T cell responses from atopic, ragweed-allergic subjects to Amb a 1, Amb a 3, Amb a 4, Amb a 5, Amb a 6, Amb a 8, Amb a 9, Amb a 10, Amb a 11, and Amb p 5 and examined their correlation with serological reactivity and sequence conservation in other allergens.

Methods: Peripheral blood mononuclear cells (PBMCs) from donors positive for IgE towards ragweed extracts after in vitro expansion for secretion of IL-5 (a representative Th2 cytokine) and IFN-γ (Th1) in response to a panel of overlapping peptides spanning the above-listed allergens were assessed.

Results: Three previously identified dominant T cell epitopes (Amb a 1 176-191, 200-215, and 344-359) were confirmed, and three novel dominant epitopes (Amb a 1 280-295, 304-319, and 320-335) were identified. Amb a 1, the dominant IgE allergen, was also the dominant T cell allergen, but dominance patterns for T cell and IgE responses for the other ragweed allergens did not correlate. Dominance for T cell responses correlated with conservation of ragweed epitopes with sequences of other well-known allergens.

Conclusions and clinical relevance: These results provide the first assessment of the hierarchy of T cell reactivity in ragweed allergens, which is distinct from that observed for IgE reactivity and influenced by T cell epitope sequence conservation. The results suggest that ragweed allergens associated with lesser IgE reactivity and significant T cell reactivity may be targeted for T cell immunotherapy, and further support the development of immunotherapies against epitopes conserved across species to generate broad reactivity against many common allergens.

Keywords: T cell epitopes; allergens; ragweed.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adolescent
  • Adult
  • Allergens / chemistry
  • Allergens / genetics*
  • Allergens / immunology*
  • Ambrosia / adverse effects*
  • Ambrosia / genetics*
  • Amino Acid Sequence
  • Antigens, Plant / chemistry
  • Antigens, Plant / genetics
  • Antigens, Plant / immunology
  • Conserved Sequence*
  • Epitopes, T-Lymphocyte / immunology
  • Female
  • Gene Expression Profiling
  • Histocompatibility Testing
  • Humans
  • Immunodominant Epitopes / chemistry
  • Immunodominant Epitopes / immunology
  • Immunoglobulin E / immunology
  • Leukocytes, Mononuclear / immunology
  • Leukocytes, Mononuclear / metabolism
  • Male
  • Middle Aged
  • Peptides / chemistry
  • Peptides / immunology
  • Plant Proteins / immunology
  • Rhinitis, Allergic, Seasonal / immunology*
  • T-Lymphocytes / immunology*
  • T-Lymphocytes / metabolism*
  • Transcriptome
  • Young Adult

Substances

  • Allergens
  • Amb a I protein, Ambrosia artemisiifolia
  • Antigens, Plant
  • Epitopes, T-Lymphocyte
  • Immunodominant Epitopes
  • Peptides
  • Plant Proteins
  • Immunoglobulin E