SHP1 Regulates Bone Mass by Directing Mesenchymal Stem Cell Differentiation

Cell Rep. 2016 Jul 19;16(3):769-80. doi: 10.1016/j.celrep.2016.06.035. Epub 2016 Jul 7.

Abstract

Osteoblasts and adipocytes are derived from a common precursor, mesenchymal stem cells (MSCs). Alterations in the normal fate of differentiating MSCs are involved in the development of obesity and osteoporosis. Here, we report that viable motheaten (me(v)) mice, which are deficient in the SH2-domain-containing phosphatase-1 (SHP1), develop osteoporosis spontaneously. Consistently, MSCs from me(v)/me(v) mice exhibit significantly reduced osteogenic potential and greatly increased adipogenic potential. When MSCs were transplanted into nude mice, SHP1-deficient MSCs resulted in diminished bone formation compared with wild-type MSCs. SHP1 was found to bind to GSK3β and suppress its kinase activity by dephosphorylating pY216, thus resulting in β-catenin stabilization. Mice, in which SHP1 was deleted in MSCs using SHP1(fl/fl)Dermo1-cre, displayed significantly decreased bone mass and increased adipose tissue. Taken together, these results suggest a possible role for SHP1 in controlling tissue homeostasis through modulation of MSC differentiation via Wnt signaling regulation.

MeSH terms

  • Adipocytes / metabolism
  • Adipogenesis / physiology
  • Animals
  • Bone Density / physiology*
  • Cell Differentiation / physiology*
  • Cells, Cultured
  • Female
  • Glycogen Synthase Kinase 3 beta / metabolism
  • Intracellular Signaling Peptides and Proteins / metabolism*
  • Male
  • Mesenchymal Stem Cells / metabolism*
  • Mesenchymal Stem Cells / physiology*
  • Mice
  • Mice, Inbred C57BL
  • Mice, Nude
  • Osteoblasts / metabolism
  • Osteoblasts / physiology
  • Osteogenesis / physiology
  • Protein Tyrosine Phosphatase, Non-Receptor Type 6 / metabolism*
  • beta Catenin / metabolism

Substances

  • Intracellular Signaling Peptides and Proteins
  • beta Catenin
  • Glycogen Synthase Kinase 3 beta
  • Protein Tyrosine Phosphatase, Non-Receptor Type 6