Autophagy Inhibits the Accumulation of Advanced Glycation End Products by Promoting Lysosomal Biogenesis and Function in the Kidney Proximal Tubules

Diabetes. 2017 May;66(5):1359-1372. doi: 10.2337/db16-0397. Epub 2017 Feb 28.

Abstract

Advanced glycation end products (AGEs) are involved in the progression of diabetic nephropathy. AGEs filtered by glomeruli or delivered from the circulation are endocytosed and degraded in the lysosomes of kidney proximal tubular epithelial cells (PTECs). Autophagy is a highly conserved degradation system that regulates intracellular homeostasis by engulfing cytoplasmic components. We have recently demonstrated that autophagic degradation of damaged lysosomes is indispensable for cellular homeostasis in some settings. In this study, we tested the hypothesis that autophagy could contribute to the degradation of AGEs in the diabetic kidney by modulating lysosomal biogenesis. Both a high-glucose and exogenous AGE overload gradually blunted autophagic flux in the cultured PTECs. AGE overload upregulated lysosomal biogenesis and function in vitro, which was inhibited in autophagy-deficient PTECs because of the impaired nuclear translocation of transcription factor EB. Consistently, streptozotocin-treated, PTEC-specific, autophagy-deficient mice failed to upregulate lysosomal biogenesis and exhibited the accumulation of AGEs in the glomeruli and renal vasculature as well as in the PTECs, along with worsened inflammation and fibrosis. These results indicate that autophagy contributes to the degradation of AGEs by the upregulation of lysosomal biogenesis and function in diabetic nephropathy. Strategies aimed at promoting lysosomal function hold promise for treating diabetic nephropathy.

MeSH terms

  • Animals
  • Autophagy / drug effects*
  • Autophagy / genetics
  • Autophagy-Related Protein 5 / genetics
  • Blood Glucose / metabolism
  • Blotting, Western
  • Cell Line
  • Cells, Cultured
  • Diabetes Mellitus, Experimental / metabolism*
  • Diabetic Nephropathies / metabolism*
  • Endocytosis
  • Epithelial Cells / drug effects*
  • Epithelial Cells / metabolism
  • ErbB Receptors / metabolism
  • Glucose / pharmacology*
  • Glycation End Products, Advanced / pharmacology*
  • Immunohistochemistry
  • Kidney Tubules, Proximal / cytology*
  • Lysosomes / drug effects*
  • Lysosomes / metabolism
  • Mice
  • Organelle Biogenesis*
  • Reverse Transcriptase Polymerase Chain Reaction

Substances

  • Atg5 protein, mouse
  • Autophagy-Related Protein 5
  • Blood Glucose
  • Glycation End Products, Advanced
  • EGFR protein, mouse
  • ErbB Receptors
  • Glucose