Formation of the IGF1R/CAV1/SRC tri-complex antagonizes TRAIL-induced apoptosis in gastric cancer cells

Cell Biol Int. 2017 Jul;41(7):749-760. doi: 10.1002/cbin.10775. Epub 2017 May 5.

Abstract

Lipid rafts provide a biological platform for apoptosis induced by tumor necrosis factor-related apoptosis-inducing ligand (TRAIL). We previously reported that insulin-like growth factor 1 receptor (IGF1R) translocation into lipid rafts helped to explain TRAIL resistance. However, it was not clear whether TRAIL resistance was caused by the interaction of IGF1R with caveolin-1 (CAV1) and the non-receptor tyrosine kinase SRC in lipid rafts of gastric cancer cells. Here, we observed high IGF1R expression in TRAIL-resistant gastric cancer cells, and showed that IGF1R combined with both CAV1 and SRC in a native complex. TRAIL was shown to promote the formation of the IGF1R/CAV1/SRC tri-complex and the activation of these three molecules. Knockdown of IGF1R or CAV1 or inhibition of SRC activity reduced the formation of this tri-complex and enhanced TRAIL-induced apoptosis. Furthermore, the overexpression of microRNA-194 reversed TRAIL resistance by reducing IGF1R expression. In summary, TRAIL increased formation of the IGF1R/CAV1/SRC tri-complex and the activation of downstream survival pathways, leading to TRAIL resistance in gastric cancer cells.

Keywords: CAV1; IGF1R; SRC; TRAIL; gastric cancer.

MeSH terms

  • Apoptosis / drug effects*
  • Caveolin 1 / metabolism*
  • Cell Line, Tumor
  • Humans
  • Membrane Microdomains / metabolism
  • MicroRNAs / metabolism
  • Mitogen-Activated Protein Kinase 3 / metabolism
  • Phosphatidylinositol 3-Kinase / metabolism
  • Receptor, IGF Type 1 / metabolism
  • Receptors, Somatomedin / metabolism*
  • Signal Transduction / drug effects
  • Stomach Neoplasms / drug therapy*
  • Stomach Neoplasms / metabolism
  • Stomach Neoplasms / pathology
  • TNF-Related Apoptosis-Inducing Ligand / antagonists & inhibitors
  • TNF-Related Apoptosis-Inducing Ligand / pharmacology*
  • src-Family Kinases / metabolism*

Substances

  • CAV1 protein, human
  • Caveolin 1
  • IGF1R protein, human
  • MIRN194 microRNA, human
  • MicroRNAs
  • Receptors, Somatomedin
  • TNF-Related Apoptosis-Inducing Ligand
  • Phosphatidylinositol 3-Kinase
  • Receptor, IGF Type 1
  • src-Family Kinases
  • Mitogen-Activated Protein Kinase 3