Nitrogen uptake and assimilation in proliferating embryogenic cultures of Norway spruce-Investigating the specific role of glutamine

PLoS One. 2017 Aug 24;12(8):e0181785. doi: 10.1371/journal.pone.0181785. eCollection 2017.

Abstract

Somatic embryogenesis is an in vitro system employed for plant propagation and the study of embryo development. Nitrogen is essential for plant growth and development and, hence, the production of healthy embryos during somatic embryogenesis. Glutamine has been shown to increase plant biomass in many in vitro applications, including somatic embryogenesis. However, several aspects of nitrogen nutrition during somatic embryogenesis remain unclear. Therefore, we investigated the uptake and assimilation of nitrogen in Norway spruce pro-embryogenic masses to elucidate some of these aspects. In our study, addition of glutamine had a more positive effect on growth than inorganic nitrogen. The nitrogen uptake appeared to be regulated, with a strong preference for glutamine; 67% of the assimilated nitrogen in the free amino acid pool originated from glutamine-nitrogen. Glutamine addition also relieved the apparently limited metabolism (as evidenced by the low concentration of free amino acids) of pro-embryogenic masses grown on inorganic nitrogen only. The unusually high alanine concentration in the presence of glutamine, suggests that alanine biosynthesis was involved in alleviating these constraints. These findings inspire further studies of nitrogen nutrition during the somatic embryogenesis process; identifying the mechanism(s) that govern glutamine enhancement of pro-embryogenic masses growth is especially important in this regard.

MeSH terms

  • Amino Acids / metabolism
  • Ammonia / metabolism
  • Biomass
  • Cell Line
  • Glutamine / metabolism*
  • Nitrogen / metabolism*
  • Picea / embryology
  • Picea / metabolism*
  • Seeds / growth & development
  • Seeds / metabolism*

Substances

  • Amino Acids
  • Glutamine
  • Ammonia
  • Nitrogen

Grants and funding

This study was conducted within the 2nd Research School of Forest Genetics, Biotechnology and Breeding (http://resschool.slu.se/), Umeå Plant Science Centre (UPSC) and Swedish University of Agricultural Sciences (SLU) Sweden, in collaboration with Svenska Skogsplantor AB (http://www.skogsplantor.se/). The work was financed by Svenska Skogsplantor AB, the Swedish Governmental Agency for Innovation Systems (VINNOVA; UPSC Berzelii Center for Forest Biotechnology 2012-01560; www2.vinnova.se; including support to U.E, T.M. and U.G.), grants awarded to T.M. (Knut & Alice Wallenberg foundation; 2011.02.12; kaw.wallenberg.org) and H.S. (Nils and Dorthi Troedsson Foundation; 859/15; www.troedssonfonden.se). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.