Role of the bicarbonate-responsive soluble adenylyl cyclase in cholangiocyte apoptosis in primary biliary cholangitis; a new hypothesis

Biochim Biophys Acta Mol Basis Dis. 2018 Apr;1864(4 Pt B):1232-1239. doi: 10.1016/j.bbadis.2017.09.022. Epub 2017 Sep 28.

Abstract

Primary biliary cholangitis (PBC) is a chronic fibrosing cholangiopathy characterized by an autoimmune stereotype and defective biliary bicarbonate secretion due to down-regulation of anion exchanger 2 (AE2). Despite the autoimmune features, immunosuppressants are ineffective while two bile acid-based therapies (ursodeoxycholic acid and obeticholic acid) have been shown to improve biochemical and histological features of cholestasis and long-term prognosis. However, the etiology and pathogenesis of PBC is largely unknown. Recently, it has been shown that microRNA-506 (miR-506) on chromosome X is up-regulated in PBC cholangiocytes and suppresses AE2 expression, which sensitizes cholangiocytes to bile salt-induced apoptosis by activating soluble adenylyl cyclase (sAC), an evolutionarily conserved bicarbonate sensor. In this review, we discuss the experimental evidence for the emerging role of the miR-506-AE2-sAC axis in PBC pathogenesis. We further hypothesize that the initial disease trigger induces an X-linked epigenetic change, leading to a female-biased activation of the miR-506-AE2-sAC axis. This article is part of a Special Issue entitled: Cholangiocytes in Health and Diseaseedited by Jesus Banales, Marco Marzioni and Peter Jansen.

Keywords: Anion exchanger 2; Apoptosis; Bile salt; Primary biliary cholangitis; Soluble adenylyl cyclase; miR-506.

Publication types

  • Review

MeSH terms

  • Adenylyl Cyclases / metabolism*
  • Apoptosis*
  • Autoimmune Diseases / etiology*
  • Autoimmune Diseases / pathology
  • Bicarbonates / metabolism
  • Bile Acids and Salts / metabolism
  • Bile Ducts / cytology
  • Bile Ducts / immunology
  • Bile Ducts / metabolism
  • Chloride-Bicarbonate Antiporters / genetics
  • Chloride-Bicarbonate Antiporters / metabolism
  • Cholangitis / etiology*
  • Cholangitis / pathology
  • Epigenesis, Genetic / genetics
  • Epigenesis, Genetic / immunology
  • Epithelial Cells / immunology
  • Epithelial Cells / metabolism
  • Epithelial Cells / pathology*
  • Female
  • Genes, X-Linked / genetics
  • Humans
  • MicroRNAs / genetics
  • MicroRNAs / metabolism
  • Signal Transduction / genetics
  • Signal Transduction / immunology
  • Up-Regulation

Substances

  • Bicarbonates
  • Bile Acids and Salts
  • Chloride-Bicarbonate Antiporters
  • MIRN506 microRNA, human
  • MicroRNAs
  • SLC4A2 protein, human
  • ADCY10 protein, human
  • Adenylyl Cyclases