Directed Differentiation of Notochord-like and Nucleus Pulposus-like Cells Using Human Pluripotent Stem Cells

Cell Rep. 2020 Feb 25;30(8):2791-2806.e5. doi: 10.1016/j.celrep.2020.01.100.

Abstract

Intervertebral disc degeneration might be amenable to stem cell therapy, but the required cells are scarce. Here, we report the development of a protocol for directed in vitro differentiation of human pluripotent stem cells (hPSCs) into notochord-like and nucleus pulposus (NP)-like cells of the disc. The first step combines enhancement of ACTIVIN/NODAL and WNT and inhibition of BMP pathways. By day 5 of differentiation, hPSC-derived cells express notochordal cell characteristic genes. After activating the TGF-β pathway for an additional 15 days, qPCR, immunostaining, and transcriptome data show that a wide array of NP markers are expressed. Transcriptomically, the in vitro-derived cells become more like in vivo adolescent human NP cells, driven by a set of influential genes enriched with motifs bound by BRACHYURY and FOXA2, consistent with an NP cell-like identity. Transplantation of these NP-like cells attenuates fibrotic changes in a rat disc injury model of disc degeneration.

Keywords: directed differentiation; human induced pluripotent stem cells; intervertebral disc disease; notochord-like cells (NCLs); nucleus pulposus (NP)-like cells.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adolescent
  • Adult
  • Animals
  • Cell Differentiation*
  • Cell Line
  • Female
  • Gene Expression Profiling
  • Genes, Reporter
  • Genome, Human
  • Green Fluorescent Proteins / metabolism
  • Human Embryonic Stem Cells / cytology
  • Humans
  • Intervertebral Disc Degeneration / pathology
  • Male
  • Notochord / cytology*
  • Nucleus Pulposus / cytology*
  • Pluripotent Stem Cells / cytology*
  • Rats, Sprague-Dawley

Substances

  • enhanced green fluorescent protein
  • Green Fluorescent Proteins