Colorectal cancer cell lines show striking diversity of their O-glycome reflecting the cellular differentiation phenotype

Cell Mol Life Sci. 2021 Jan;78(1):337-350. doi: 10.1007/s00018-020-03504-z. Epub 2020 Mar 31.

Abstract

Alterations in protein glycosylation in colorectal cancer (CRC) have been extensively studied using cell lines as models. However, little is known about their O-glycome and the differences in glycan biosynthesis in different cell types. To provide a better understanding of the variation in O-glycosylation phenotypes and their association with other molecular features, an in-depth O-glycosylation analysis of 26 different CRC cell lines was performed. The released O-glycans were analysed on porous graphitized carbon nano-liquid chromatography system coupled to a mass spectrometer via electrospray ionization (PGC-nano-LC-ESI-MS/MS) allowing isomeric separation as well as in-depth structural characterization. Associations between the observed glycan phenotypes with previously reported cell line transcriptome signatures were examined by canonical correlation analysis. Striking differences are observed between the O-glycomes of 26 CRC cell lines. Unsupervized principal component analysis reveals a separation between well-differentiated colon-like and undifferentiated cell lines. Colon-like cell lines are characterized by a prevalence of I-branched and sialyl Lewis x/a epitope carrying glycans, while most undifferentiated cell lines show absence of Lewis epitope expression resulting in dominance of truncated α2,6-core sialylated glycans. Moreover, the expression of glycan signatures associates with the expression of glycosyltransferases that are involved in their biosynthesis, providing a deeper insight into the regulation of glycan biosynthesis in different cell types. This untargeted in-depth screening of cell line O-glycomes paves the way for future studies exploring the role of glycosylation in CRC development and drug response leading to discovery of novel targets for the development of anti-cancer antibodies.

Keywords: Cell lines; Colorectal cancer; Glycomics; Mass spectrometry; O-Glycosylation; Porous graphitized carbon liquid chromatography.

MeSH terms

  • Carbohydrate Sequence
  • Cell Differentiation*
  • Cell Line, Tumor
  • Chromatography, High Pressure Liquid
  • Colorectal Neoplasms / metabolism
  • Colorectal Neoplasms / pathology
  • Glycomics / methods*
  • Glycosylation
  • Glycosyltransferases / genetics
  • Glycosyltransferases / metabolism
  • Humans
  • Phenotype
  • Polysaccharides / analysis*
  • Polysaccharides / metabolism
  • Principal Component Analysis
  • Tandem Mass Spectrometry

Substances

  • Polysaccharides
  • Glycosyltransferases